Clitobius, Mulsant & Rey, 1859

Purchart, Luboš & Kamiński, Marcin Jan, 2017, A taxonomic review of the genus Clitobius with description of a new species from Oman (Coleoptera: Tenebrionidae), Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 57, pp. 139-163 : 151-152

Key to the species of Clitobius

1. Metathoracic wings absent or reduced (e.g., Fig. 3D). ................................................. 2

– Metathoracic wings fully developed. ............................................................................ 5

2. Elytral intervals flat (Fig. 3E). ....................................................................................... 3

– Elytral intervals convex ( Fig. 2F ). ................................................................................ 4

3. Body size: 6.2–7.0 mm. Sides of pronotal disc with sparsely punctured distinct submarginal groove or depression along the lateral edge. ... Clitobius grandis Fairmaire, 1896

– Body size ca. 5.0 mm. Sides of pronotal disc without submarginal groove or depression; surface with coarse punctuation; in some cases lateral margination slightly widened near the posterior ends. ........................................... Clitobius strongyloides Fairmaire, 1892

4. Body size: 5.0– 5.5 mm. Metathoracic wings absent. Hypomera covered with extremely fine punctures. Elytra black and shiny. Elytral rows with coarse punctures (distributed densely than those on pronotal disc). .................... Clitobius cribricollis ( Allard, 1882)

– Body size: 6.6–7.0 mm. Metathoracic wings reduced but present (Fig. 3D). Hypomera covered with dense and coarse punctures. Elytra dull with greenish metallic shimmer. Elytral rows with fine and elongated punctures (distributed similarly to those on pronotal disc). ................................................................ Clitobius obesus ( Waterhouse, 1881)

5. Pronotum widest directly at base. Elytral humeri strongly protruding outwards. ........... ............................................................................................. Clitobius kochi Ferrer, 1995

– Pronotum widest in the middle or near base (Fig. 3G). Elytral humeri not protruding outwards (Fig. 3G). ....................................................................................................... 6

6. Protibiae equipped with a prominent apical tooth. Male protarsi widened (see FERRER 2001). .................................................................. Clitobius endroediorum Ferrer, 2001

– Protibiae slender, without a prominent apical tooth. Male protarsi not widened. ......... 7

7. Dorsal side of the body covered with coarse and noticeable setae, arranged on each elytral interval in three parallel rows (Fig. 3I). Posterior pronotal angles rounded (Fig. 3G). ............................................................................ Clitobius ovatus ( Erichson, 1843)

– Dorsal side of the body glabrous or covered with fine setae (Fig. 3H). Posterior pronotal angles rounded or sharp (e.g., Fig. 3H). ........................................................................ 8

8. Body elongated (body length to maximum width ratio = 2.0–2.3). Antennomere III relatively short (length ratio antennomeres II / III ca. 1.4). Antennomeres evenly widened, not forming well designated club. Posterior pronotal angles sharp (Fig. 3H). In some cases, pronotal disc with a longitudinal midline without punctures. ............................... .................................................................... Clitobius oblongiusculus ( Fairmaire, 1875)

– Body oval (body length to maximum width ratio = 1.6–1.9). Antennomere III relatively elongated (length ratio antennomeres II / III ca. 2.1). Five terminal antennomeres forming a club ( Fig. 2E ). Posterior pronotal angles rounded ( Fig. 1B ). Centre of pronotal disc evenly covered with punctures. ................................. Clitobius omanicus sp. nov.

Figures

Fig. 2. Detailed morphology of Clitobius obesus (Waterhouse, 1881) (A–C, F) and C. omanicus sp. nov. (D, E, G). Abbreviations: a – trochanter base; b – femur base; c – rounded posterior angle of pronotal disc; d – not depressed sides of pronotal disc; e – convex intervals; f – ventral; g – dorsal; h – lateral view.

Fig. 1. Clitobius obesus (Waterhouse, 1881) (A) and C. omanicus sp. nov. (B). Habitus (dorsal view) and aedeagal tegmen (lateral view).