Inocellia

Liu, Xingyue, Aspöck, Horst, Zhang, Weiwei & Aspöck, Ulrike, 2012, New species of the snakefly genus Inocellia Schneider, 1843 (Raphidioptera: Inocelliidae) from Yunnan, China, Zootaxa 3298, pp. 43-52 : 44-45

Key to males of Inocellia

1. Wing membrane hyaline (except pterostigma), immaculate..................................................... 2

- Wing membrane distinctly brown apically.............................. I. elegans Liu, H. Aspöck, Yang & U. Aspöck

2. Gonocoxite 9 wider than long ( Fig. 8 )………….( I. crassicornis group)........................................... 3

- Gonocoxite 9 more or less produced posteriad, slightly or distinctly longer than wide ( Fig. 12 ; Liu et al. 2010b: figs. 17, 28; U. Aspöck et al. 2011: fig. 45)………….( I. fulvostigmata group)................................................. 10

3. Gonarcus (fused gonocoxites 11) with paired or unpaired protrusions posteriad ( Liu et al. 2010a: fig. 31)............... 4

- Gonarcus without posterior protrusion..................................................................... 6

4. Posterior protrusion of gonarcus bifurcate ( Liu et al. 2010a: fig. 32)............................................. 5

- Posterior protrusion of gonarcus paired, slender and digitiform ( Liu et al. 2010a: fig. 26)........................................................................................... I. digitiformis Liu, H. Aspöck, Yang & U. Aspöck

5. Gonocoxite 9 with a feebly prominent stylus on inner side ( Liu et al. 2010a: fig. 30)..................... I. fujiana Yang

- Gonocoxite 9 with a long and acutely pointed stylus on inner side (U. Aspöck et al. 2009: fig. 5)........................................................................................... I. shinohara U. Aspöck, Liu & H. Aspöck

6. Endophallus proximally with a pair of processes dorsolaterally ( Liu et al. 2010a: fig. 6, see arrow)..................... 7

- Endophallus proximally without process ( Liu et al. 2010a: fig. 15)............................................... 8

7. Endophallus distally with a pair of tufts of bristles (H. Aspöck et al. 1991: fig. 2499)................ I. japonica Okamoto

- Endophallus distally with only two bristles or none ( Liu et al. 2010a: fig. 6)..................................................................................................... I. biprocessus Liu, H. Aspöck, Yang & U. Aspöck

8. Gonocoxite 9 with rows of bundles anterior to stylus (H. Aspöck et al. 1991: fig. 2502)... I. taiwana H. Aspöck & U. Aspöck

- Gonocoxite 9 without tuft anterior to stylus, but sometimes with a rather small process posterior to stylus................ 9

9. Gonarcus subtrapezoidal in caudal view ( Liu et al. 2010a: fig. 17); endophallus with two pairs of bristle tufts ( Liu et al. 2010a: fig. 15)......................................................................... I. crassicornis (Schummel)

- Gonarcus subquadrate in caudal view ( Fig. 9 ); endophallus with only one pair of tufts on dorsal surface ( Fig. 9 ).................................................................................................... I. nigra sp. nov.

10. Gonocoxite 9 with digitiform or unguiform stylus........................................................... 11

- Gonocoxite 9 with rather small or broadly inflated and roundly tapered stylus..................................... 16

11. Gonocoxite 9 in caudal view with a rounded lobe ventrally ( Fig. 15 ); gonarcus medially domed, but without conspicuous pro- trusion posteriad ( Fig. 16 ).............................................................................. 12

- Gonocoxite 9 in caudal view without ventral lobe ( Liu et al. 2010b: fig. 20); gonarcus medially with conspicuous protrusion posteriad ( Liu et al. 2010b: fig. 22)...................................................................... 14

12. Endophallus without bristles (H. Aspöck et al. 1991: fig. 2518)............... I. bhutana H. Aspöck, U. Aspöck & Rausch

- Endophallus with several bristles and tufts................................................................. 13

13. Gonocoxite 9 with a slender digitiform stylus (U. Aspöck et al. 2011: fig. 6); endophallus with only one pair of tufts on dorsal surfaces (U. Aspöck et al. 2011: fig. 3)............................. I. longispina U. Aspöck, Liu, Rausch & H. Aspöck

- Gonocoxite 9 with an unguiform stylus ( Fig. 15 ); endophallus with five tufts on both dorsal and ventral surface ( Figs. 14, 16 ).................................................................................... I. yunnanica sp. nov.

14. Gonocoxite 9 with an additional process anterior to stylus (U. Aspöck et al. 2011: fig. 47); stylus thickly digitate, bearing sev- eral bristles (U. Aspöck et al. 2011: fig. 47)........................... I. cornuta U. Aspöck, Liu, Rausch & H. Aspöck

- Gonocoxite 9 without additional process anterior to stylus; stylus unguiform, without bristles........................ 15

15. Gonocoxite 9 with a tuft anterior to stylus on inner side ( Liu et al. 2010b: fig. 21); stylus with a geniculate claw, curved dorsad ( Liu et al. 2010b: fig. 20)........................................... I. hamata Liu, H. Aspöck, Yang & U. Aspöck

- Gonocoxite 9 without tuft anterior to stylus on inner side ( Liu et al. 2010b: fig. 39); stylus simply unguiform, slightly curved ventrad ( Liu et al. 2010b: fig. 38)............................................................ I. sinensis Navás

16. Gonocoxite 9 with stylus inconspicuous, rather small, and bearing no bristle or tuft on inner side ( Liu et al. 2010b: fig. 7)...................................................................... I. cheni Liu, H. Aspöck, Yang & U. Aspöck

- Gonocoxite 9 with prominent stylus, broadly inflated, and bearing a tuft on inner side.............................. 17

17. Gonocoxite 9 with stylus bilobed (U. Aspöck et al. 2011: fig. 13).......... I. bilobata U. Aspöck, Liu, Rausch & H. Aspöck

- Gonocoxite 9 with stylus not bilobed...................................................................... 18

18. Gonarcus medially without protrusion ( Liu et al. 2010b: fig. 32)...... I. obtusangularis Liu, H. Aspöck, Yang & U. Aspöck

- Gonarcus medially with conspicuous protrusions............................................................ 19

19. Stylus of gonocoxite 9 nearly cone-like (U. Aspöck et al. 2011: fig. 32); gonarcus medially with a distally bifurcated protrusion and with ventral corners acutely angulated (U. Aspöck et al. 2011: fig. 35)... I. striata U. Aspöck, Liu, Rausch & H. Aspöck

- Stylus of gonocoxite 9 lobe-like; gonarcus medially with a pair of protrusions and with ventral corners rounded (H. Aspöck et al. 1991: figs. 2508, 2515, 2516)………. I. fulvostigmata U. Aspöck & H. Aspöck. ................................ 20

20. Pterostigma yellow......................................... I. fulvostigmata fulvostigmata U. Aspöck & H. Aspöck

- Pterostigma brown................................... I. fulvostigmata nigrostigmata H. Aspöck, U. Aspöck & Rausch

Figures

FIGURES 8 – 11. Inocellia nigra sp. nov. 8. Male genital segments, lateral view; 9. Male genital segments, caudal view; 10. Male genital segments showing internal structures, lateral view; 11. Fused parameres (complex of fused gonocoxites, gonapophyses, and gonostyli 10), dorsal view. e: ectoproct; ep: endophallus; g: gonarcus (fused gonocoxites 11); gx 9: gonocoxite 9; p: fused parameres; ps: pseudostylus (basal parts of gonapophyses 9); s: stylus or stylus-like process (gonostylus 9); S 8, 9: sternite 8 and 9; T 8, 9: tergite 8 and 9. Scale bar = 0.5 mm.

FIGURES 12 – 19. Inocellia yunnanica sp. nov. 12. Male genital segments, lateral view; 13. Male genital segments, dorsal view; 14. Male genital segments, ventral view; 15. Male genital segments, caudal view; 16. Male genital segments showing internal structures, lateral view; 17. Gonarcus (fused gonocoxites 11), dorsal view; 18. Fused parameres (complex of fused gonocoxites, gonapophyses, and gonostyli 10), dorsal view; 19. Hypandrium internum, ventral view. Scale bar = 0.5 mm.