Dyscinetus dubius (Olivier, 1789)

Moore, Matthew R., Cave, Ronald D. & Branham, Marc A., 2018, Annotated catalog and bibliography of the cyclocephaline scarab beetles (Coleoptera, Scarabaeidae, Dynastinae, Cyclocephalini), ZooKeys 745, pp. 101-378 : 229-230

Dyscinetus dubius (Olivier, 1789)

Melolontha dubia Olivier, 1789: 32 [original combination].

Dyscinetus dubius (Olivier) [new combination by Bates 1891: 32].

Dyscinetus (Palechus) dubius (Olivier) [new subgeneric classification by Casey 1915: 174].

Dyscinetus dubius (Olivier) [removal of subgeneric classification by Arrow 1937b: 17].

syn. Geotrupes lugubris Quensel, 1806: 21 [original combination].

Melolontha geminata Fabricius [synonymy by Schönherr 1817: 187].

syn. Dyscinetus frater Bates, 1888: 312 [original combination].

Dyscinetus dubius (Olivier) [synonymy by Endrődi 1966: 387].

syn. Dyscinetus (Dyscinetus) obtusus Casey, 1915: 170 [original combination].

Dyscinetus frater Bates [synonymy by Chapin 1932: 295].

syn. Melolontha geminata Fabricius, 1801: 166-167 [original combination].

Chalepus geminatus (Fabricius) [new combination by MacLeay 1819: 149-150].

Cyclocephala geminata (Fabricius) [new combination by Laporte 1840: 124].

Chalepus geminatus (Fabricius) [revised combination by Burmeister 1847: 78-79].

Dyscinetus geminatus (Fabricius) [new combination by Harold 1869a: 123].

Dyscinetus dubius (Olivier) [synonymy by Prell 1937a: 187].

Types.

Type material of M. dubia is apparently missing ( Endrődi 1966). Lectotype ♂ of M. geminata deposited at ZMUK, now housed at ZMUC ( Endrődi 1966). Endrődi (1966) did not find the type material of G. lugubris . Lectotype ♂ of D. frater at BMNH ( Endrődi 1966). Type of D. obtusus at USNM ( Endrődi 1966).

Distribution.

ARGENTINA: Buenos Aires, Entre Ríos, Misiones. BELIZE: Cayo. BOLIVIA: Beni, Chuquisaca, Cochabamba, La Paz, Santa Cruz. BRAZIL: Acre, Amazonas, Bahia, Espírito Santo, Goiás, Mato Grosso, Pará, Paraná, Pernambuco, Rio de Janeiro, Rio Grande do Norte, Rio Grande do Sul, Rondônia, Santa Catarina, São Paulo. CHILE. COLOMBIA: Amazonas, Antioquia, Atlántico, Boyacá, Caquetá, Cauca, Casanare, Chocó, Córdoba, Cundinamarca, Huila, Meta, Risaralda, Santander, Tolima, Valle del Cauca. COSTA RICA: Alajuela, Cartago, Guanacaste, Heredia, Limón, Puntarenas, San José. CUBA: Camagüey, Ciego de Ávila, Isla de la Juventud, Matanzas, Pinar del Río, Sancti Spíritus. ECUADOR: Guayas. EL SALVADOR: Ahuachapán, Cabañas, Cuscatlán, La Libertad, San Miguel, San Salvador, Santa Ana. FRENCH GUIANA: Cayenne, St.-Laurent du Maroni. GUATEMALA: Alta Verapaz, Baja Verapaz, Chiquimula, Escuintla, Guatemala, Huehuetenango, Izabal, Petén, Quetzaltenango, Retalhuleu, San Marcos, Santa Rosa, Suchitepéquez, Zacapa. GUYANA: Demerara-Mahaica, Mahaica-Berbice. HONDURAS: Atlántida, Choluteca, Mayagua, Cortés, El Paraíso, Francisco Morazán, Gracias a Dios, La Paz, Olancho. MEXICO: Campeche, Chiapas, Distrito Federal, Guerrero, Jalisco, Nayarit, Oaxaca, Quintana Roo, San Luis Potosí, Sinaloa, Tabasco, Tamaulipas, Tlaxcala, Veracruz. NICARAGUA: Carazo, Granada, Managua, Masaya, RAA Sur, Río San Juan, Rivas. PANAMA: Bocas del Toro, Panama Canal Zone, Chiriquí, Coclé, Colón, Darien, Panamá, San Blas. PARAGUAY: Central, Cordillera, Guaíra, Paraguarí. PERU. SURINAME: Paramaribo District. TRINIDAD AND TOBAGO: Trinidad. VENEZUELA: Amazonas, Apure, Aragua, Bolívar, Carabobo, Delta Amacuro, Falcón, Guárico, Mérida, Monagas, Portuguesa, Táchira, Trujillo, Zulia.

References.

Olivier 1789, Fabricius 1801, Quensel 1806, Schönherr 1817, MacLeay 1819, Dejean 1821, 1833, 1836b, Latreille 1829, Laporte 1840, Sturm 1843, Burmeister 1847, Erichson 1848a, Reiche 1859, Fauvel 1861, Harold 1869a, Bates 1888, 1891, Ohaus 1900, Casey 1915, Cleare 1925, 1930, Chapin 1932, Squire 1932, 1933, Dash 1934, Prell 1937a, Arrow 1937b, Anonymous 1900, 1940, Blackwelder 1944, Guimarães 1944, Wiehe 1951, Figueroa-P. 1952, Lima 1953, Roze 1955, Gibson and Carrillo 1959, Van Dinther 1960, Zimsen 1964, Endrődi 1966, 1973a, 1985a, Dechambre 1979a, Morón 1979, Remillet 1988, Aguilera et al. 1993, Thomas 1993, Lobo and Morón 1993, Maes 1987, 1994, Ratcliffe and Morón 1997, Sanchez Soto 1998, Andreazze and Fonseca 1998, Ratcliffe 1986, 2002a, 2003, Restrepo-Giraldo et al. 2003, Pardo-Locarno et al. 1995, 2005a, Riehs 2005, Fernández García 2006, Ratcliffe and Cave 2006, Pacheco-F. et al. 2008, Marques and Gil-Santana 2009, Gasca-Álvarez and Amat-García 2010, Joly and Escalona 2010, Ponchel 2011, Neita-Moreno and Yepes 2011, Neita-Moreno 2011, Krajcik 2005, 2012, Breeschoten et al. 2013, Pardo-Locarno 2013, Otavo et al. 2013, García-López et al. 2013, Ratcliffe et al. 2013, Deloya et al. 2014a, Ratcliffe et al. 2013, 2015, Ratcliffe and Cave 2015, García-Atencia and Martínez-Hernández 2015, López-García et al. 2015, Albuquerque et al. 2016.