Carabus (Archiplectes) besleticus Kurnakov, 1972

Solodovnikov, Igor A., Zamotajlov, Alexandr S., Fominykh, Dmitriy D. & Titarenko, Andrey Y., 2014, Contribution to the knowledge of the Carabus (Archiplectes) satyrus Kurnakov, 1962, species complex in Abkhazia (Coleoptera, Carabidae, Carabini), ZooKeys 463, pp. 21-56 : 24-25

Carabus (Archiplectes) besleticus Kurnakov, 1972 stat. n. Figs 1, 2, 3, 4, 5, 6, 7, 16-21, 22-27, 28-33, 34-41, 42-49, 50-57, 58-61, 62-65, 66-73, 74-77

Remarks.

The structure of the endophallus testifies to the attribution of this form to the reitteri group. Saccellus large, conical, smoothly rounded apically, with faintly developed or missing S-shaped fold on the right, paracellar lobe very faintly developed, endophallus with rather symmetric position of inner structures (when viewed frontally). Ostium lobe large, of dulled conical shape. Lamella of aedeagus elongated, with prominent hollow on the right before apex. Aggonoporius large, strongly sclerotized. This type of endophallus is characteristic for all subspecies described subsequently, earlier attributed to Carabus (Archiplectes) satyrus . Based on such pronounced differences in the structure of the endophallus from that of Carabus (Archiplectes) satyrus , it is proposed to raise the status of Carabus (Archiplectes) besleticus from subspecies to species.