Megalopsenella pouilloni, Jouault & Ngô-Muller & Pouillon & Nel, 2021

Jouault, Corentin, Ngô-Muller, Valérie, Pouillon, Jean-Marc & Nel, André, 2021, New Burmese amber fossils clarify the evolution of bethylid wasps (Hymenoptera: Chrysidoidea), Zoological Journal of the Linnean Society 191 : -

MEGALOPSENELLA POUILLONI SP. NOV.

( FIGS 5, 6)

urn:lsid:zoobank.org:act:578342A1-78A5-4F5E-9A08-E5D3E1B46A83 .

Holotype: Female , accession number Po 55 (complete and well-preserved female specimen in a rounded piece of amber measuring 20 × 20 × 5 mm, without syninclusions), housed in the Musée d’Histoire Naturelle de Colmar, France.

Type locality and horizon: Noije Bum Hill, Hukawng Valley, Kachin State, Myanmar; Lower Cenomanian, Upper Cretaceous.

Etymology: Named after Jean-Marc Pouillon, who donated the holotype.

Description: Female; body about 7 mm long, not depressed, almost smooth and not strongly foveolate; LFW 4.31 mm; castaneous; wings setose. LH 1.13 mm; WH 0.29 mm; WF 1.3 mm; HE 0.33 mm; OOL 0.55 mm; WOT 0.32 mm; DAO 0.14 mm; VOL 0.18 mm. Head not pubescent, smooth; mandible elongate, teeth not visible; six maxillary palpomeres, three (?) visible labial palpomeres; gena slightly projecting forward; reduced clypeus medially emarginate; antennal sockets partially closed dorsally by a tooth-shaped projection; antenna with 11 cylindrical flagellomeres; scape 0.47 × 0.22 mm, pedicel 0.1 2 × 0.1 2 mm, F1 0.1 4 × 0.1 2 mm (all measurements refer to length × width), F2 0.15 × 0.13 mm, F3 0.15 × 0.13 mm, F4 0.15 × 0.13 mm, F5 0.15 × 0.11 mm, F6 0.18 × 0.11 mm, F7 0.15 × 0.11 mm, F8 0.16 × 0.10 mm, F9 0.17 × 0.10 mm, F10 0.17 × 0.09 mm; F11 0.3 × 0.09 mm; occipital carina strongly marked; compound eye without microsetae; ocellar triangle not visible. Mesosoma with dorsum smooth, not sculptured;

KEY TO GENERA OF † PROTOPRISTOCERINAE 1. Antenna with 13 antennomeres with short scape; forewing venation usually developed but sometimes reduced...............................................................................................................................................................2

- Antenna with 12 antennomeres, scape elongated; forewing venation always reduced (Cenomanian: Myanmar) ................................................................................................................... Cretapristocera gen. nov.

2. Head elongated with small eyes .......................................................................................................................3

- Head almost globose with large eyes ...............................................................................................................4

3. Mandibles only with few teeth (five) along masticatory margin; legs with pro- and meta-femora clearly thickened; forewing cell 2 R 1 open (Cenomanian: Myanmar) ........................... Bethylitella Cockerell, 1917a

- Mandibles with small teeth along masticatory margin and with basal one elongated: legs, except pro- femora, only slightly thickened; forewing cell 2 R 1 almost closed (Eocene: Baltic amber) .............................. ................................................................................................................................ Protopristocera Brues, 1923

4. Notauli well developed, parapsidal furrow weak (Eocene: Oise amber) .... Gynopteron Falières & Nel, 2019

- Notauli weak, parapsidal furrow well-visible (Eocene: Baltic amber) ................. Bethylopteron Brues, 1933

Forewing covered with microtrichia and bordered with small setae; anterior border not angularly incurved anterior to pterostigma; seven closed cells present, and all veins C, Sc + R, Rs, M + Cu, Cu, A tubular; pterostigma rather broad; poststigmal abscissa of R1 long; hindwing 3.23 mm long with five hamuli. Metasoma c. 5.76 mm long, with short petiole; length of clearly visible tergites: T1 1.45 mm, T2 1 mm, T3 at least 0.8 mm; short part of sting visible.

femora strongly thickened; tibia thin, pretarsal claws with strong subapical tooth; tibial spur formula 1-1-2; notauli weakly marked; parapsidal furrows very weak; mesoscutellum smooth; metapectal–propodeal complex flat, rectangular, not foveolate, and with no posterior spines; posterior margin of propodeum straight.