Geostiba (Tropogastrosipalia) biformis, Homson, 1974

Homson, 1974, Lichenology in North America, 1947 - 1972., Linzer biologische Beiträge 61, pp. 45-55: 45-55

publication ID

http://doi.org/10.5281/zenodo.5434894

persistent identifier

http://treatment.plazi.org/id/6B4D2217-FFDC-AC62-F15E-209BFDDEFBD3

treatment provided by

Valdenar

scientific name

Geostiba (Tropogastrosipalia) biformis
status

sp.n.

Geostiba (Tropogastrosipalia) biformis   sp.n. ( Figs 54-63 View Figs 54-62 View Fig , Map 3 View Map 3 )

Holotype: TR [20] - Mu ÷la, 15 km ENE Mu ÷la, 1190 m, pasture with stones, 37°14'58N, 28°30'07E, 10.IV.2006, V. Assing GoogleMaps   / Holotypus Geostiba biformis   sp.n. det. V. Assing 2006 (cAss). Paratypes: 5, 4: same data as holotype (cAss, OÖLL) GoogleMaps   ; 2, 1: same data, but leg Wunderle (cWun) GoogleMaps   ; 3, 2: TR [21] - Denizli, 60 km E Mu ÷la, S Kale, 1280 m, 37°25'39N, 28°53'18E, 11.IV.2006, V. Assing (cAss) GoogleMaps   ; 6, 10: same data, but leg. Wunderle (cWun, cAss) GoogleMaps   ; 1: TR [22] - Denizli, 60 km E Mu ÷la, S Kale, 1240 m, 37°23'12N, 28°53'41E, 11.IV.2006, V. Assing (cAss) GoogleMaps   .

D e s c r i p t i o n: Small species, 2.2-3.0 mm (abdomen fully extended). Coloration: head dark brown; pronotum and elytra yellowish red to brown; abdomen blackish, with the apex and often also the anterior segments paler; legs testaceous; antennae yellowish brown to reddish brown.

Facies as in Fig. 54 View Figs 54-62 . Head and pronotum with variable, shallow to distinct microreticula-

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tion ( Fig. 55 View Figs 54-62 ). Eyes relatively small ( Fig. 56 View Figs 54-62 ), weakly protruding from lateral outline of head, approximately 1/3 the length of postocular region in dorsal view or even smaller.

Pronotum with moderately pronounced sexual dimorphism. Elytra with weak sexual dimorphism, less than 0.6 times as long as pronotum.

Abdomen with distinct, but shallow microsculpture, with fine and moderately sparse puncturation; posterior margin of tergite VII with or without very narrow rudiment of a palisade fringe.

(large): pronotum approximately as long as wide or weakly oblong, 1.15-1.20 times as wide as head; posterior margin of pronotum in the middle broadly and weakly concave, postero-laterally sinuate ( Fig. 55 View Figs 54-62 ); elytra with moderately dense and somewhat granulose puncturation, with distinct extensive impression, elevated (somewhat bulging) lateral margins, and with narrow and weakly elevated sutural carina extending along anterior 2/3 of suture ( Figs 55, 57 View Figs 54-62 ); abdominal tergites III-IV with ( Fig. 58 View Figs 54-62 ) or without indistinct tubercles; process of tergite VII short and stout ( Figs 59-60 View Figs 54-62 ); posterior margins of tergite and sternite VIII broadly convex; median lobe of aedeagus with slender cristal process ( Fig. 61 View Figs 54-62 ); apical lobe of paramere shaped as in other species of the subgenus.

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: pronotum approximately 1.10 times as wide as head and about as wide as long, posterior margin broadly convex; elytra unmodified, without carina, at most with very shallow impressions, and with fine puncturation; posterior margins of tergite and sternite VIII weakly convex; spermatheca as in Fig. 62.E t y m o l o g y View Figs 54-62 : The name (Lat., adj.) alludes to the dimorphism of the male abdominal tergites III-IV (see comments on intraspecific variation).

I n t r a s p e c i f i c v a r i a t i o n: As in other species of the subgenus, the male secondary sexual characters are subject to considerable intraspecific variation. In smaller males, the pronotum may be almost unmodified, the elytral puncturation may be weakly granulose, the elytral impression are often very shallow, and the sutural carinae and the process of the tergite VII may be completely absent.

Remarkably, the two sampled populations, one (1) from the area to the northeast of Mu÷la and the other (2) from the area approximately 60 km to the east of Mu÷la, are distinguished by a constant character. The abdominal tergites III-IV of the males from population 2 each have a minute median tubercle ( Fig. 58 View Figs 54-62 ), which is absent in the males from population 1. However, since no additional distinguishing characters were found and since these populations are not separated by deep valleys, this difference is here attributed to intra- (possibly clinal) rather than interspecific variation.

C o m p a r a t i v e n o t e s: Using the key in ASSING (2005a), G. biformis   would key out at couplets (67-71). In order to account for the new species, as well as for G.

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aydinica sp.n. and G. solodovnikovi   sp.n., the key is modified as follows:

67: abdominal tergites III and IV each with smooth subcircular tubercle near anterior impression. Species from western, southern or eastern Anatolia. ( G. biformis   , in which tergites III and IV may or may not have indistinct tubercles, will key out in both alternatives.).....................................................................................67a

-: abdominal tergites III and IV unmodified. Species from northern or western Turkey, or from Konya ...............................................................................................69

67a: tubercles on tergites III and IV minute, almost indistinct ( Fig. 58 View Figs 54-62 ); process of tergite VII apically rounded in antero-dorsal view ( Figs 59-60 View Figs 54-62 ); median lobe of aedeagus as in Fig. 61 View Figs 54-62 . Western Anatolia (Mu÷la, W-Denizli) ( Map 3 View Map 3 )......................... ............................................................................................................. G. biformis   sp.n.

-: tubercles on tergites III and IV distinct; process of tergite VII apically acute in antero-dorsal view. Eastern and southern Anatolia.....................................................68

68 Pronotum with more pronounced microreticulation.: elytra with wider sutural carinae (Fig. A00a: 10); aedeagus larger and with more slender cristal process (Figs A00a: 11-12). Southern Anatolia (Antalya)............................ G. brachati ASSING  

- Pronotum with rather shallow microreticulation.: aedeagus smaller and with stouter cristal process. Eastern and northeastern Anatolia........................................68a

68a Eyes smaller ( Fig. 21 View Figs 20-29 ), approximately one third the length of postocular region in dorsal view.: posterior margin of pronotum usually broadly concave, margins between this concavity and posterior angles straight, i. e. pronotum posteriorly gradually tapering ( Fig. 22 View Figs 20-29 ); tubercles on tergites III-IV less pronounced and not more shining than surrounding area; process of tergite VII more slender, with almost parallel sides, and apically rounded in antero-dorsal view ( Fig. 25 View Figs 20-29 ); median lobe of aedeagus with slightly stouter cristal process ( Fig. 27 View Figs 20-29 ). Northeastern Anatolia (Erzurum: Mescit Da÷ları) ( Map 4 View Map 4 ).............................. G. solodovnikovi   sp.n.

- Eyes larger, approximately half the length of postocular region in dorsal view or nearly so (Fig. A01a: 44).: posterior margin of pronotum indistinctly concave to smoothly convex, margins between this concavity and posterior angles sinuate, i. e. pronotum posteriorly more abruptly tapering (Fig. A01a: 44); tubercles on tergites III-IV more pronounced and with somewhat more shine than surrounding area; process of tergite VII broad-based and apically acute (i. e. of triangular shape) in antero-dorsal view ( Fig. 43 View Figs 36-43 in Assing 2001a); median lobe of aedeagus with more slender cristal process (Figs A01a: 40-41). Eastern Anatolia (Bitlis: surroundings of Tatvan)...................................................................................... G. bitlisensis ASSING  

69: pronotum (in large!) distinctly oblong, more or less extensively depressed, strongly projecting posteriorly, and with broadly and distinctly concave hind margin ( Fig. P83b View Figs 78-84 : 1); elytra with distinctly elevated (bulging) lateral margins; process of tergite VII wide-based, apically rounded, and in lateral view rather slender; aedeagus: Figs. P83b View Figs 78-84 : 2-3. Southern Anatolia (Konya)....... G. iconiensis PACE  

-: lateral margins of elytra not distinctly elevated. Species from northern and western Turkey (Thrace, northern Anatolia)...............................................................70

70: pronotum (in large!) distinctly oblong and strongly projecting posteriorly, its posterior margin in the middle narrowly concave (Fig. A00b: 1); elytra with relatively long sutural carinae; process of tergite VII in antero-dorsal view slender and apically rounded, in lateral view more massive (Figs. A00b: 3-4); aedeagus with cristal process of median lobe of characteristic shape (Figs. A00b: 2, P83b: 25-26). Northern Anatolia (Kastamonu) ................................. G. kastamonuensis PACE  

-: pronotum often less oblong, its posterior margin broadly concave or truncate; aedeagus with cristal process much smaller or of different shape.............................70a

70a: pronotum (even in large) usually about as long as wide, at most 1.05 times as long as wide, its posterior margin in the middle weakly concave and laterally sinuate ( Fig. 55 View Figs 54-62 ); elytra with weakly pronounced (i. e. weakly elevated) sutural carina ( Figs 55, 57 View Figs 54-62 ); median lobe of aedeagus as in Fig. 61 View Figs 54-62 . West Anatolia: Mu÷la, W-Denizli............................................................................................. G. biformis   sp.n.

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-: pronotum in large moderately to distinctly oblong and of different shape; elytra with more pronounced sutural carina; median lobe of aedeagus with cristal process of different shape .........................................................................................70b

70b: pronotum (in large) more oblong (up to 1.25 times as long as wide) and more strongly projecting posteriorly ( Fig. 46 View Figs 45-52 ); elytra with more strongly elevated and longer sutural carina and with coarser and more distinctly granulose puncturation ( Figs 46, 48 View Figs 45-52 ); process of tergite VII and median lobe of aedeagus as in Figs 49-51 View Figs 45-52 . West Anatolia (Aydın): Aydın Da ÷ları ......................... G. aydinica   sp.n   .

-: pronotum less oblong and less strongly projecting posteriorly; elytra with less pronounced sutural carina and with finer and less distinctly granulose puncturation. Species from Thrace and northeastern Anatolia..........................................................71

71: pronotum (in large) slightly more strongly projecting posteriorly and with weakly concave lateral margins near posterior angles (Fig. A01a: 14); abdomen less finely punctate; process of tergite VII wide-based and apically rounded or obtuse (Fig. A01a: 19); cristal process of aedeagus much longer and larger (Figs. A01a: 12-13). Northeastern Anatolia (Artvin)............................. G. artvinensis ASSING  

-: pronotum in large less strongly projecting posteriorly, lateral margins near posterior angles not distinctly concave, hind margin in the middle truncate or broadly concave; abdomen very finely punctate; process of tergite VII more slender and apically more acute; aedeagus with cristal process of median lobe very short and thin. Thrace (surroundings of Istanbul) .................... G. turcica (BERNHAUER)  

D i s t r i b u t i o n a n d b i o n o m i c s: Geostiba biformis   was recorded from three localities, one in Mu÷la province, in the vicinity of Mu÷la, and two in the west of Denizli province, some 60 km to the east of Mu÷la ( Map 3 View Map 3 ). The type specimens were found under stones and sifted from grass roots in pastures on calcareous soil ( Fig. 63 View Fig ) at altitudes of 1190-1280 m.

V

Royal British Columbia Museum - Herbarium