Batrachomatus Clark, 1863

Hendrich, Lars & Balke, Michael, 2013, Revision of Australian Matini diving beetles based on morphological and molecular data (Coleoptera, Dytiscidae, Matinae), with description of a new species, ZooKeys 293, pp. 41-64 : 43

Batrachomatus Clark, 1863

Batrachomatus Clark, 1863: 15 (type species Batrachomatus wingii Clark, 1863, by monotypy); Zimmermann 1919: 215 (cat.); Mouchamps 1964: 137 (descr.); Watts 1978: 119 (descr.); Watts 1985: 23 (cat.); Lawrence et al. 1987: 351 (cat.); Nilsson 2001: 261 (cat.); Larson 1993: 50 (cat.); Watts 2002: 31, 46 (cat.).

Allomatus Mouchamps, 1964: 137 (type species Allomatus wilsoni Mouchamps, 1964, by original designation); Watts 1978: 117 (descr.); Watts 1985: 23 (cat.); Lawrence et al. 1987: 351 (cat.); Nilsson 2001: 261 (cat.); Watts 2002: 30, 46 (cat.); syn. n.

Remarks.

Medium-sized (TL = 6.9-9.6 mm), elongate, shiny and flattened mainly black diving beetles, unicolorous or with testaceous or ferruginous markings on elytra. Epipleuron in apical half more than twice as wide as base of longer spur of metatibia. Both parameres abruptly narrowed near middle. Outer metatarsal claw curved, prosternum with median furrow. Body covered with dense fine punctures and/or very fine microreticulation, meshes irregular, polygonal.

Allomatus was justified as a genus based only on the presence of a reticulated pronotum and elytra without any punctuation in contrast to Batrachomatus which has a non-reticulate but densely and finely punctate pronotum and elytra ( Mouchamps 1964, Watts 1978). Brachomatus larsoni sp. n. has characters from both genera: a polygonal double reticulation with punctures at the intersections of all meshes.

The cox1 sequence data show (Fig. 15) that Allomatus wilsoni is sister to Batrachomatus daemeli , Batrachomatus wingi sister to these two, and Allomatus nannup sister to all of these. We use Matus bicarinatus (Say, 1823) from North America as well as Hygrobia maculata Britton, 1981 and Hygrobia wattsi Hendrich, 2001 as outgroups ( Hygrobia were pruned for Fig. 15). Consequently Allomatus Mouchamps, 1964 is here synonymised with Batrachomatus Clark, 1863.