Asymmetricata humeralis ( Walker 1858 ) Ho, 2019

Ho, - Z., 2019, The Luciolinae of S. E. Asia and the Australopacific region: a revisionary checklist (Coleoptera: Lampyridae) including description of three new genera and 13 new species, Zootaxa 4687 (1), pp. 1-174 : 52-55

publication ID

https://doi.org/ 10.11646/zootaxa.4687.1.1

publication LSID

lsid:zoobank.org:pub:CE73264D-C234-4B82-A634-CAD6254C5957

DOI

https://doi.org/10.5281/zenodo.4688817

persistent identifier

https://treatment.plazi.org/id/6C3DA91C-5108-1860-FF0E-FCD3EE0D1D20

treatment provided by

Plazi

scientific name

Asymmetricata humeralis ( Walker 1858 )
status

comb. nov.

Asymmetricata humeralis ( Walker 1858) View in CoL comb. nov.

Figs 154−174 View FIGURES 154–164 View FIGURES 165−174

Colophotia humeralis Walker 1858: 282 .

Type. Colophotia humeralis . Two syntype males. CEYLON. Male 1 labelled 1. Male symbol (handwritten); 2. Ceylon; 69 106 on back of label (circular blue label handwritten); 3. Humeralis (handwritten on white label); Colophotia humeralis Walker A.N.H. (type) (handwritten on blue label). Male 2 labelled 1. Male symbol (handwritten); 2. Ceylon (handwritten on blue label); 3. Colophotia humeralis Walker A.N. H. type (handwritten on blue label). ( NHML). Figs 154, 155, 158, 159, 160 View FIGURES 154–164 .

Luciola humeralis (Walker) View in CoL . Olivier 1902: 80. Gorham 1903: 327. McDermott 1966: 105.

Luciola doriae Olivier 1885: 358 View in CoL ; 1902: 78. McDermott 1966: 103. New synonymy.

Type. Luciola doriae . Two syntype males. CEYLON. Not examined by these authors. ( MCSN).

Luciola impressa Olivier 1910b: 532 View in CoL . McDermott 1966: 106. New synonymy.

Type. Luciola impressa . Holotype male, 2 syntype females. CEYLON . Holotype male Labelled 1. Type (printed on red label); 2. Andrewes bequest (printed); 3. Kandy Ceylon (printed) 5–09 (handwritten); 4. 2497 (handwritten); 5. Luciola impressa Ern Oliv. ♂ ♀ (handwritten on pink label). Syntype female 1 labelled: 1. Kandy Ceylon 5–09 (green label); 2. 2497 (hand- written ink); 3. Andrewes bequest (printed). Syntype female 2 labelled 1. Peradeniya Ceylon 5–09; 2. 2498; 3. Andrewes bequest. ( NHML). Figs 165−167, 171−174 View FIGURES 165−174 .

Luciola notatipennis Olivier 1909a: 318 View in CoL . New synonymy.

Type. Luciola notatipennis . Possible syntype male. INDIA. Labelled 1. Nilgiri Hills (printed); 2. 824 (handwritten); 3. Andrewes bequest (printed); 4. Prob. Oliv. Luciola doriae var. notatipennis Ern. Oliv. (handwritten on pink label may be in Olivier’s handwriting). ( NHML).

Other specimens examined. S INDIA. Nilgiri Hills Singara P. S. Nathan male. S. Malabar 8.96 Walayr forest P. S. Nathan female ( PRAGUE). Mormugao Goa, July 1925 JC Bridwell 29 males, 3 females. ( USNM) .

Diagnosis. Dorsal colouration differing from that of As. circumdata and As. ovalis in having paler brown elytra with darker brown markings restricted to base and apex; elytra of impressa type brownish with narrowly paler lateral margin and very narrowly pale suture and only elytral bases darker brown ( Fig. 165 View FIGURES 165−174 ); pronotum either entirely pale yellow (in humeralis type Fig. 154 View FIGURES 154–164 , Nilgiri Hills and S. Malabar specimens), with paired median brown spots (in impressa type Fig. 167 View FIGURES 165−174 ), or completely brownish (in notatipennis syntype Fig. 157 View FIGURES 154–164 ). Differing from As. circumdata in having bipartite LOs in V7 ( Fig. 158 View FIGURES 154–164 ). Apart from the dorsal colouration As. humeralis comb. nov. is distinguished most obviously by the aedeagal sheath tergite which is emarginated on its right side, and the distal apex of the sheath sternite which is obliquely truncate ( Figs 161, 162 View FIGURES 154–164 ).

Remarks. The synonymy established here was suggested by Olivier as a note in his collection at MNHN (LB examination 2013). The emarginations of abdominal T8 (emarginate on its inner right side) and that of the sheath tergite (emarginate on its inner left side) suggest an easier egress for the aedeagus dorsally.

We now know of four species having a pale yellowish or pale brown dorsum with darker markings at base and apex. Atyphella telokdalam sp. nov. is known only from Sumatra, while Aty. palauensis is from the Republic of Palau. Luciola flavescens (Boisduval) from ‘Oceanie’ is included in species incertae sedis.

Wijekoon et al. (2016) confirmed the occurrence of both L. humeralis and L. impressa in Sri Lanka and recognised them both as belonging to Asymmetricata . They also recorded L. doriae but made no comment about its possible position.

USNM

USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]

NHML

Natural History Museum, Tripoli

MCSN

Museo Civico di Storia Naturale, Verona

USNM

Smithsonian Institution, National Museum of Natural History

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Lampyridae

SubFamily

Luciolinae

Genus

Asymmetricata

Loc

Asymmetricata humeralis ( Walker 1858 )

Ho, - Z. 2019
2019
Loc

Luciola impressa

McDermott, F. A. 1966: 106
Olivier, E. 1910: 532
1910
Loc

Luciola notatipennis

Olivier, E. 1909: 318
1909
Loc

Luciola humeralis

McDermott, F. A. 1966: 105
Gorham, H. S. 1903: 327
Olivier, E. 1902: 80
1902
Loc

Luciola doriae

McDermott, F. A. 1966: 103
Olivier, E. 1902: 78
Olivier, E. 1885: 358
1885
Loc

Colophotia humeralis

Walker, F. 1858: 282
1858
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