Paratorchus tilli, Irmler, 2015

Irmler, Ulrich, 2015, New Neotropical genera and species of the tribe Osoriini (Coleoptera: Staphylinidae: Osoriinae), Beiträge Zur Entomologie = Contributions to Entomology 65 (1), pp. 173-196 : 185-186

Paratorchus tilli spec. nov.

( Figs 14A–C , 18A , 19C )

Type material: Holotype male: Mexico, Veracruz, 1 mi S Huatusco , 1344 m elevation, collected by Berlese, 02.08.1969, leg. S. & J. Peck ( AMNH).

Paratypes: Two females with same data as holotype ( AMNH, UIC) .

Etymology: The specific name of this small species derived from the name of my little grand son Till.

Description: Length: 2.8 mm. Colouration: light reddish brown; legs and antennae lighter brown.

Head: 0.35 mm long, 0.49 mm wide; without eyes; fore-head approximately semicircular; anterior edge of clypeus even; punctation moderately dense and deep; on average, interstices between punctures at least as wide as diameter of punctures; on disc two times as wide as 1. Blind, abdomen not pubescent ................................................................................................................. P. tilli spec. nov.

- At least with few omatidia .................................................................................................................................................. 2

2. Abdomen pubescent ........................................................................................................................................................... 3

- Abdomen not pubescent .................................................................................................................................................... 6

3. Sides of pronotum evenly narrowed from middle to posterior angles, posterior angles obtuse ................................. ................................................................................................................. P. brasiliensis ( BERNHAUER & SCHUBERT, 1910)

- Sides of pronotum nearly parallel in posterior half ........................................................................................................ 4 diameter; laterally with few setae; setae extremely short; without microsculpture; between normal punctures with sparse micro-punctation; temples with deep netlike microsculpture.

4. Large species of at least 3.0 mm length, sides of pronotum parallel, not sinuate in front of posterior angles ....... 5

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Small species of 2.5 least, sides of pronotum at least slightly sinuate in front of posterior angles .............................. ................................................................................................................................................... P. panamae ( IRMLER, 2005) 5. Sides of pronotum evenly rounded, pronotal punctation deeper and denser, interstices between punctures as wide as diameter of punctures ..................................................................................................... P. bolivianus ( IRMLER, 2005)

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Sides of pronotum slightly sinuate in middle, pronotal punctation finer and sparser, interstices between punctures much wider than diameter of punctures ....................................................................... P. columbiensis ( IRMLER, 1987) 6. Large species of at least 4.0 mm length ............................................................................... P. breviatus ( IRMLER, 2007)

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Smaller species of 2.5 to 2.8 mm length ........................................................................................................................... 7 7. Pronotum deeply and densely punctate, surface polished without microsculpture, shiny .......................................... ........................................................................................................................................... P. chiriquensis ( IRMLER, 2005)

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Pronotum finely and sparsely punctate, surface with netlike microsculpture, matt ............ P. chilensis ( IRMLER, 2005)

Antennae as long as head and half of pronotum combined; first antennomere thick; thicker than second oval and conical third antennomere; second and third antennomere equal in length and longer than following antennomeres; antennomeres four to eleven increasing in width; fourth longer than wide; tenth nearly twice as wide as long; surface polished and shiny.

Pronotum: 0.51 mm long, 0.64 mm wide; widest in anterior third; slightly narrowed to anterior angles in convex curve; in posterior two third evenly narrowed to posterior angles; nearly parallel; posterior angles rectangular; lateral margin distinct; in dorsal aspect visible throughout its total length; at posterior angles with indistinct impression; punctation deeper and denser than on head; on average, interstices between punctures as wide as diameter of punctures; between coarse punctures with sparse micro-punctation; with irregular microsculpture; surface less shiny than on head.

Elytra: 0.37 mm long, 0.67 mm wide; sides slightly widened to posterior angles; shoulders rectangular; posterior edge slightly emarginate; punctation still deeper and coarser than on pronotum; partly coriaceous; with coriaceous ground-sculpture; surface less shiny than on pronotum.

Abdomen densely and deeply punctate; similar as on pronotum; dorsally punctures without setae; laterally and ventrally with short yellow setae; surface without microsculpture; polished and shiny.

Aedeagus elongate; apical lobe smoothly rounded ending in acute apex; paramera long and sinuate; longer than apical lobe; endophallus with long and straight basal part and short spiral part at apex.

Diagnosis: The species can be easily identified by the absence of eyes. All other species in the genus have at least few omatidia. The overall habitus resembles that of P. panamae , but P. panamae has at least 5 omatidia and the pronotum is more densely punctate.

Figures

Fig. 10–14: Tumboecus struyvei (10); Osoriosetus hanagarthi (11), Neosorius brasiliensis (12), N. mexicanus (13), Paratrochus tilli (14); antenna (A), protibia in posterior aspect (B), aedeagus (C); antenna (A), protibia in posterior aspect (B), aedeagus (C); scale bar: 0.1 mm.

Fig. 18: Head, pronotum, and elytra of Paratrochus tilli (A), P. brasiliensis (B), P.chilensis (C), P. columbiensis (D), P. panamae (E), and P. chiriquensis (F); scale bar: 0.5 mm.

Fig. 19: Head, pronotum, and elytra of Paratrochus bolivianus (A), P. breviatus (B), and head in lateral aspect of P. tilli (C), P.chilensis (D), P. panamae (E), P. chiriquensis (F), P. columbiensis (G), P. brasiliensis (H), P. bolivianus (I), and P. breviatus(K); scale bar of A and B: 0.5 mm; scale bar of C–K: 0.2 mm.