Tambana indeterminata, Behounek, G., Han, H. L. & Kononenko, V. S., 2015

Behounek, G., Han, H. L. & Kononenko, V. S., 2015, A revision of the genus Ta mb ana Moore, 1882 with description of eight new species and one subspecies (Lepidoptera, Noctuidae: Pantheinae). Revision of Pantheinae, contribution XIII, Zootaxa 4048 (3), pp. 301-351 : 323-324

Tambana indeterminata sp. n.

( Figs 57–60 , 74–78 , 112, 113 )

Type material. Holotype. INDIA: male, Prov. Arunachal-Pradesh, Distr., road Dirang Bomdila, 2450 m, 27°266′N, 92°427′E, Bretschneider leg., 3–5.viii.2006, slide GB7403 male, ZSM Lep 48884, coll. GB/ ZSM; Paratypes: 1 male, same data, slide GB12164 male, coll. JS; CHINA: 1 male, Prov. Yunnan, Mekong river valley, local collector leg. 14.v.2000, slide GB12145 male, coll. WS; 1 male, Prov. Xizang, Han Mi- Mo Tuo, 2100 m, 18– 27.viii.2005, Huang, Zhou, Tang leg., genit. slide HHL-1773; 1 male, Xizang, Linzhi, 108 km, 19.viii.2011, Zh. Pan leg., genit. slide HHL-2601; 1 male, same region and collector, 80 km, 24.vii.2011, genit. slide HHL-2601S, all coll. NEFU; THAILAND: 1 male, Prov. Chiang Mai, 1100 m, 4 km SE of Pang, Faen, 13.i.2004, Attila Szabó leg., coll. AB.

Note. The specimen labelled as BC ZSM Lep 48884 have been used for barcoding. Molecular distance based on the Kimura two-parameter model for COI DNA barcodes between T. indeterminata and the nearest species T. annamica is 4,44%. Barcode difference between T. indeterminata and T. glauca is 5,28%.

Diagnosis. Externally T. indeterminata is very similar to T. glauca , but differs by somewhat bigger size, darker and duller yellowish-brown with metallic shining ground colour of forewing, less developed greyish dusting, less dentate and more contrast whitish transversal lines and by deep yellow with dark black-brownish terminal band hindwing. The new species well separated from T. glauca by the differences in their DNA barcodes.

Description. Adult ( Figs 57–60 , 74–78 ). Wingspan 36–38 mm. Head, tegulae and patagia yellowish-white, patagia bordered with black, thorax dark yellowish-grey with brown crest; ground colour of forewing dark yellowish-brown with metallic shining, much darker than in T. glauca ; basal field black, with white bordering; subbasal field dark yellowish-brown; antemedial line black, less waved than in T. glauca , with thin white inner borderline; medial field dark yellowish-brown with diffused medial shadow expressed as black patches in costal and ventral part of wing; orbicular as small orange-yellowish dot on black background; cell between stigmata filled with black; orbicular small, like yellow-brown or brown streak encircled with white line; small orange or yellowish dash lies from reniform towards postmedial line; postmedial line black, less dentate than in T. glauca , outlined inwardly by white contrast borderline; forming prominent white mark in costal area above reniform; subterminal field rather wide, dark yellowish-brown, with metallic shining; subterminal line less dentate and less contrast than in T. glauca , marked by bordering with brown terminal field, which forms clear whitish-brown triangular mark; between two extensions of subterminal line; terminal line as row of thin black streaks, bordered with thin orange line; cilia dark yellowish-brown, whitish opposite veins. Hindwing with deep yellow basal part and dark broad brownish-black terminal band; cilia grey-brown, pale.

Male genitalia ( Fig. 112, 113 ). Male genitalia very similar to T. glauca , but slightly differ by narrower valva, higher juxta, shorter and more straight harpe, narrower aedeagus and more developed subapical fields of cornuti. Uncus short, thinner than in T. glauca , curved; tegumen broad, shorter than vinculum, with prominent penicular extensions; vinculum broadly V-shaped; transtilla with broad leave-like lobes; juxta shield-like, broad and higher the in T. glauca ; valva relatively short, somewhat narrower than in T. glauca , with parallel margins, sacculus narrow, elongate; in apical third ventral margin of valva rounded and gradually constricted distally, apex rounded; harpe lies in distal third of valva, straight, relatively short. Aedeagus relatively short narrower than in T. glauca , carina sclerotised; vesica broadly-tubular, rather short, bearing large basal diverticulum covered with field of small spines and large apical diverticulum with two patches of small dense cornuti more developed than in T. glauca .

Female unknown.

Etymology. The specific name is given by the difficulty in identification of the species and separation it from T. glauca .

Distribution and bionomy. North India (Arunachal Pradesh), Southwest China (Provs. Yunnan: Mekong valley; Xizang=Tibet), North Thailand (Chiang Mai). The species occurs in secondary and primary forest at elevation 2000–2400 m. Moths fly in May and August.

Figures

FIGURES 57 – 64. Tambana spp., adults. 57. T. indeterminata sp. n., male, North India, holotype, slide GB 7403 (GB / ZSM); 58. T. indeterminata sp. n., male, paratype, China, Yunnan, slide GB 12145 (WS / ZSM); 59. T. indeterminata sp. n., male, paratype, China, Xizang, slide HHL- 1773 (NEFU); 60. T. indeterminata sp. n., male, paratype, North India, slide GB 12164 (JS); 61. T. annamica, male sp. n., holotype, North Vietnam, slide GB 7405 (GB / ZSM); 62. T. annamica, male sp. n., paratype, North Vietnam, slide GB 7406 (GB / ZSM); 63. T. annamica sp. n., male, paratype, North Vietnam, slide RL 6669 (HNHM); 64. T. annamica sp. n., male, paratype, North Vietnam (AB / ZSM).

FIGURES 65 – 88. Tambana spp., adults. 65. T. annamica stumpfi ssp. n., male, holotype, China, Guangxi, slide GB 12113 (PGM); 66. T. annamica stumpfi ssp. n., male, paratype, China, Shaanxi (JS); 67. T. annamica stumpfi ssp. n., male, paratype, China, Guangxi, slide GB 12112 (PGM); 68. T. annamica stumpfi ssp. n., male, paratype China, Guangdong, slide HHL- 1500 - 1 (NEFU); 69 – 73. Series of T. glauca sp. n. (North India, North Vietnam, North Thailand, Myanmar, Southwest China); 74 – 78. Series of T. indeterminata sp. n. (North India, North Thailand, Southwest China); 79 – 83. Series of T. annamica sp. n. (North Vietnam); 84 – 88. Series of T. annamica stumpfi ssp. n. (China: Shaanxi, Guangxi, Guangdong).

FIGURES 110 – 113. Tambana spp., male genitalia. 110. T. glauca, North Vietnam, holotype of behouneki, slide 5375 (WS / ZFMK); 111. T. glauca, Thailand, slide GB 7423 (GB / ZFMK); 112. T. indeterminata sp. n., holotype, North India, slide GB 7403 (GB / ZSM); 113. T. indeterminata sp. n., paratype, China, Guangxi slide HHL- 2606 - 17403 (NEFU).