Tambana naumanni Speidel & Kononenko, 1998

Behounek, G., Han, H. L. & Kononenko, V. S., 2015, A revision of the genus Ta mb ana Moore, 1882 with description of eight new species and one subspecies (Lepidoptera, Noctuidae: Pantheinae). Revision of Pantheinae, contribution XIII, Zootaxa 4048 (3), pp. 301-351 : 308-309

Tambana naumanni Speidel & Kononenko, 1998

( Figs 13, 14 , 93 , 123 )

Tambana naumanni Speidel & Kononenko, 1998 , Esperiana , 6: 554, Pl. T: 18, U: 19. Type-locality: China, Prov. Fujian, Kuatun, 2300 m. Holotype: male, coll. ZFMK.

Material examined. Types: Holotype, male, Guadun (2300 m) 27°40'N 117°40'E (Fukien) J.K. Klapperich, 21.vi.1938, coll ZFMK and series of paratypes from China (same locality, 21.iv, 23.iv, 30.iv, 1.v, 18.v, 19.v.1938, coll. ZFMK and from Vietnam, coll. WS coll. HNHM, slides VK1784 male ( ZFMK); WS412 ( ZFMK); MH10931 ( HNHM). Other material examined. CHINA: 1 male, Prov. Jiangxi, border Jiangxi-Fujian, Wuy Shan, 50 km SO Yingtan, 1600 m, 27°950 N, 117°400 E, Siniaev V. leg., slide VK 07- 090311 male, iv–v.2002, coll. AB; 1 male, same data, coll. GB/ ZSM; 1 male; 1 female, same data, slide GB8529 male ( BC ZSM Lep 58638) coll. GB/ ZSM; 1 female, Prov. Jiangxi, Wuyi Shan, Xipiahe, 1500 m, 27°55′N, 117°20′E, Sinjaev V. & loc. collector leg., 1– 30.vi.2003, slide GB12121 female, coll. PG; 1 male; Prov. Guangxi, Dayao Shan, Liuzhou, 100 km SO, Umg. Jingxiu, 1200 m, 23°45′N, 109°45′E, Sinyaev V. & Team, leg., 16–30.iii.2005, slide GB12119 male, coll. PG; Prov. Shaanxi, Taibei Shan, Tsinling Mts, [Qin Ling], 1900 m, 33°53′N, 107°49′E, Siniaev V. leg., ix.2004, slide GB12168 male, coll. JS.

Note. The specimen labelled as BC ZSM Lep 58638 has been used for barcoding. Molecular distance based on the Kimura two-parameter model for COI DNA barcodes between T. naumanni to the nearest species T. helmuti is 5,91%; the distance between T. naumanni and T. burmana is 6,26%.

Diagnosis. Adult ( Figs 13, 14 ). Wingspan 38–46 mm. Compared with T. burmana and T. plumbea the species differs by biserrate antennae, more elongate and more acute forewing shape, less developed silver-shining suffusion and details of wing pattern, forewing colouration with pale whitish-yellow central part, well defined brownish-grey terminal band and basal area.

Male genitalia ( Fig. 93 ). Compared with T. burmana and T. plumbea the male genitalia differ by broad valva with parallel margins, short vinculum and shape and arming of vesica, which formed by tree patches of dense small cornuti and setae. Uncus rather broad, short; tegumen broad, equal in length to vinculum, without penicular extensions; vinculum U-shaped; transtilla with relatively narrow branches; juxta shield-like, broad, valva moderate long, with parallel margins, slightly curved having oblique distal margin; bearing a patch of strong setae on the top, harpe moderate long, almost straight, slightly extend distal margin of valva; pointed apically. Aedeagus moderate in length, straight, slightly extended apically; carina short, sclerotised, scobinate; vesica tubular, recurved, with subapical diverticulum, bearing medial and subapical patches of small cornuti.

Female genitalia ( Fig. 123 ). Compared with related T. burmana and T. plumbea the female genitalia have rather broad antrum, short ductus bursae and ribbed sclerotised cervix bursae. Ovipositor quadrangular, papillae anales moderate broad, quadrangular; antrum triangular, weakly sclerotised; ductus bursae short, membranous; corpus bursae sack-like, with sclerotised patch in joining with ductus bursae.

Distribution and bionomy. Northwest and East China (Provs Shaanxi, Jiangxi, Fujian), North Vietnam. The species occurs in low and meso-montane forest at elevation 1000–1500 m. Moths fly in March, April, May, June, July, September.

Figures

FIGURES 9 – 16. Tambana spp., adults. 9. T. burmana, male, holotype, Myanmar (NHRM); 10. T. burmana, male, China, Fujian (PGM); 11. T. burmana, female, China, Sichuan, slide GB 12122 (PGM); 12. T. burmana, female, China, Sichuan, slide GB 12130 (PGM); 13. T. naumanni, male, holotype, China, Guangdong (ZFMK); 14. T. naumanni, male, paratype, China, Guangdong (ZFMK); 15. T. plumbea, female, holotype, Japan, Tokyo (NHM); 16. T. plumbea, male, Russia, Sakhalin (GR).

FIGURES 92 – 94. Tambana spp., male genitalia. 92. T. burmana, paratype, Myanmar, slide VK / NHRM- 2 (NHRM); 93. T. naumanni, paratype, China, Guangdong, slide VK 1784 (ZFMK); 94. T. plumbea, Russia, Sakhalin, slide VK 277 (IBSS).

FIGURES 118 – 123. Tambana spp., female genitalia. 118. T. albiplaga, North Vietnam, slide GB 8254 (GB / ZSM); 119. T. arctoides, Myanmar, slide GB 12161 (AB); 120. T. bella, China, Yunnan, slide GB 12117 (PGM); 121. T. fansipana sp. n., holotype, North Vietnam, slide GB 7402 (GB / ZSM); 122. T. burmana, China, Sichuan, slide GB 12122 (PGM); 123. T. naumanni, paratype, China, Guangdong, WS 412 (ZFMK).