Aoplonema princeps (Uhler)

Forero, D., 2008, Revision And Phylogenetic Analysis Of The Hadronema Group (Miridae: Orthotylinae: Orthotylini), With Descriptions Of New Genera And New Species, And Comments On The Neotropical Genus Tupimiris, Bulletin of the American Museum of Natural History 2008 (312), pp. 1-172 : 33-38

Aoplonema princeps (Uhler) Figures 1A–F, 4–6, 8–12, 14

Hadronema princeps Uhler, 1894: 251 [n. sp.]; Knight, 1925: 182 [list]; Kelton, 1959: 30 [male genitalia]; Kelton, 1980: 225 [diagnosis, hosts, distribution].

Hadronema (Aoplonema) princeps: Knight, 1928: 177 [subgeneric placement]; Carvalho, 1958: 68 [catalog]; Henry and Wheeler, 1988: 410 [catalog].

Hadronema uhleri Van Duzee, 1928: 182 [n. sp.]. NEW SYNONYMY.

Hadronema (Aoplonema) uhleri: Knight, 1928: 181 [key, distribution]; Knight, 1968: 94 [distribution]; Henry and Wheeler, 1988: 410 [catalog].

Hadronema (Aoplonema) uniformis Knight, 1928: 180 [n. sp.]; Knight, 1968: 95 [diagnosis, distribution]. NEW SYNONYMY.

Hadronema uniforme: Steyskal, 1973: 207 [emendation].

Hadronema (Aoplonema) uniforme: Henry and Wheeler, 1988: 410 [catalog].

Aoplonema princeps: Kerzhner and Schuh, 1995: 4 [new status]; Schuh, 1995: 81 [catalog]; Maw et al., 2000: 116 [list].

Aoplonema uhleri: Kerzhner and Schuh, 1995: 4 [new status]; Schuh, 1995: 81 [catalog], Maw et al., 2000: 116 [list].

Aoplonema uniformis: Kerzhner and Schuh, 1995: 4 [new status]; Schuh, 1995: 81 [catalog].

DIAGNOSIS: Recognized by the anteocular portion of the head about half as long as total head length (fig. 5); the small eyes that do not reach the dorsal margin of the head in lateral view (fig. 5); and the weakly elevated and well-defined calli on pronotum.

Aoplonema princeps resembles A. rubrum due to the similar structure of the vesica (fig. 10), but it is easily distinguished from A. rubrum by the structure of the head, small size of eyes (fig. 5), and comparatively larger interocular space (table 1). Aoplonema princeps is easily distinguished from A. nigrum by the structure of the male genitalia, in particular the larger size of the genital capsule, and longer rami of the vesica (fig. 10). Females of all species are difficult to identify, and only tentative placements can be done in association with males.

REDESCRIPTION: Male: From delicate to nearly robust species, total length 3.56–5.10. COLORATION (fig. 1): HEAD: Reddish black to completely black; sometimes mandibular and maxillary plates red, line at base of plates connecting antennal bases with eyes pigmented black; labrum and labium dark brown, sometimes segment I of labium weakly reddish; antennae dark brown. THO- RAX: Collar, pronotum, mesoscutum, and scutellum reddish black (fig. 1A), red (fig. 1D, E), orange (fig. 1F), sometimes anterior lobe of pronotum nearly black (e.g., fig. 1B, F), sometimes mesoscutum and scutellum dark red (fig. 1B); thoracic pleura red, orange, or nearly black; venter black. Hemelytra (fig. 1): Dark brown to black (fig. 1D, F); sometimes areas next to claval suture whitish (fig. 1A); cuneus reddish brown to red with anterior portion white (fig. 1A), or completely dark brown (fig. 1D), or weakly reddish black (fig. 1B, D). Legs: Coxae, trochanters, and femora red (fig. 1D, E), reddish black (fig. 1A, B), or orange (fig. 1F), sometimes anterior femora with distal half paler (fig. 1B), sometimes irregular dark spots on surface; tibiae and tarsi dark brown. ABDOMEN: Reddish black. GENITALIA: Genital capsule reddish black or black; proctiger pale red or brown; parameres brown. STRUC- TURE: HEAD: Gently declivent; anteocular region long, about one-third of head length (figs. 4A, 5); frons rounded; gula usually shorter than buccula; eyes small, not reaching dorsal margin of head in lateral view, interocular distance large (fig. 5; table 1). THORAX: Calli on pronotum weakly elevated and strongly impressed. Hemelytra: Nearly parallel; cuneus from nearly flat to weakly deflexed, about as long as wide (fig. 1). GENITALIA: Genital capsule relatively large (fig. 6); parameres as in figure 6; left portion of vesica compressed dorsoventrally at apex, strongly denticulate, rami long, almost reaching apex of right portion (fig. 10); right portion strongly curved upward, sometimes weakly sinuate apically, directed to the right, variously denticulate, from a single acute apex to weakly expanded and denticulate (fig. 10).

Female: Similar to male, weakly larger, ovoid, total length 4.26–4.95. COLORA- TION: Similar to male (fig. 1). ABDOMEN: Sternites reddish black, sternite IX sometimes from bright red to reddish black. STRUC- TURE: GENITALIA: Interramal sclerites and dorsal lobe of interramal sclerite as in figure 8, sigmoid process barely denticulate; sclerotized rings ovoid, elongate, posterior margin not produced, medial margin projecting cephalad, accessory sclerite not enlarged, rounded with a small denticulate at apex (fig. 8); anterior wall and inner margins of first gonapophyses as in figure 9.

DISTRIBUTION: Widely distributed from the 100th meridian to the west, and from southern Canada to Baja California ( Mexico) (fig. 14).

HOSTS: Aoplonema princeps exhibits a large and diverse set of plant associations, among them with Asteraceae , Cupressaceae , Fabaceae , Fagaceae , Lamiaceae , Pinaceae , Rhamnaceae , Rosaceae , and Sarcobataceae species (see appendix 1). In addition to these plants, one specimen was found preying upon ‘‘ A. pomi ’’ ( Aphis pomi ?) (AMNH_PBI 00107173), which is not surprising because many phytophagous Miridae , including some Orthotylini , are known to feed on other insects ( Wheeler, 2001). Young (1984b) mentioned that A. princeps (as H. princeps , H. uhleri , and H. uniformis ) was associated with cantharidin traps. I have examined Aoplonema specimens associated with cantharidin traps, and I was able to confirm that they are A. princeps .

DISCUSSION: VARIABILITY: Aoplonema princeps , as delimited here, is a morphologically variable species (see generic discussion). Aoplonema uhleri and A. uniforme are considered synonyms of A. princeps because they cannot be distinguished by any combination of characters. Several color variants are thus present in the broadly conceived A. princeps (fig. 1A–F).

Aoplonema princeps was described by Uhler (1894) as very similar to the reddish H. robusta Uhler (5 Lopidea robusta ), but narrower and having a distinct whitish cuneus (fig. 1A). Females of this variant usually have the sternite IX reddish black (fig. 1A, arrow). In addition to the whitish cuneus, some specimens of this color variant have the claval suture whitish (e.g., fig. 1A). Some specimens associated with Sarcobatus have a wide whitish claval suture, whereas specimens from other hosts have a small or no whitish fascia at all. Even specimens from the same collection event (i.e., same locality, same date), the width of this fascia varies (e.g., AMNH_PBI 00107077, AMNH_PBI 00107070). The white extent of the cuneus is also variable, not only among localities but within the same collection event, varying from a distinct transverse white band on the anterior margin of the cuneus to almost nonexistent, in which case the cuneus is nearly completely red (e.g., AMNH_PBI 00102501, AMNH_PBI 00102502).

Colorado specimens are usually darker on the legs and hemelytra with the cuneus slightly reddish (fig. 1B, arrow). Females of this variant usually have sternite IX bright red (fig. 1B, arrow). One examined female (AMNH_PBI 00107139) is as robust as other Colorado specimens, but it has sternite IX reddish black and a small whitish anterior marking on the cuneus, making it an intermediate between the white cuneus color variant and the darker Colorado variant. Some females of the white cuneus color variant may exhibit a reddish sternite IX, looking similar to females from Colorado (e.g., fig. 1C, arrow). Some specimens from Colorado, although robust and with sternite IX bright red, have a small white band on the cuneus (e.g., AMNH_PBI 00107201–AMNH_ PBI 00107203).

The slender body variant, with mostly orange femora, and dark brown cuneus without white markings (fig. 1F, arrow), is what Knight (1928) described as A. uniforme . This variant feed mostly on Sarcobatus (see appendix 1), but it may be associated with other plants as well. Intermediate forms between this variant and the more robust variant with red femora from Colorado exist. For instance, a robust female with dark orange legs (AMNH_PBI 00104806) is intermediate in color between these two. A male with red femora resembles the orange leg variant due to the narrow body (AMNH_ PBI 00105586). Some specimens from Utah that have a clear white band on the cuneus are narrow, and the femora are orange, being intermediate between this orange leg color variant and the white cuneus one (AMNH_ PBI 00102408–AMNH_PBI 00102412).

The variant with red femora, dark cuneus, and robust body is what Van Duzee (1928) described as A. uhleri (fig. 1D). The uniform dark cuneus is also present in the orange leg variant, although in the red femora variant the cuneus may also be reddish (fig. 1D, E, arrows) resembling the Colorado variant. A specimen from New Mexico (AMH_PBI 00105613) is similar to the coloration of A. uhleri paratypes, but it is darker on the hemelytra, middle, and posterior legs, and is barely more robust, being intermediate between the Colorado variant and those paratypes.

The distribution of these population color variants is indicated in figure 14. Because many specimens are intermediate between these populations, as exemplified above, I decided to arbitrarily assign whether a particular specimen belonged to either variant in those cases.

Vesica structure in A. princeps is quite variable and not correlated with coloration patterns or geographical distribution. For example, one A. uhleri paratype (AMNH_ PBI 00077861) (figs. 1D, 10) and one A. uniforme paratype (AMNH_PBI 00104807) (figs. 1F, 10) share almost identical vesical morphology, although coloration patterns are quite different. Similarly, specimens from the Colorado population usually have the right portion of the vesica weakly expanded and nearly flattened (fig. 10, AMNH_PBI 00102778, AMNH_PBI 00102782), although specimens from Wyoming and California, with a distinctive whitish cuneus, have a similar vesica structure (fig. 10, AMNH_ PBI 00079863, AMNH_PBI 00107301).

All aforementioned examples support the hypothesis of a single taxon that can be distinguished from other Aoplonema species , although being variable in coloration and size. The variability exhibited by A. princeps is similar to what is found in Lopidea nigridea Uhler, 1895 , another Orthotylini occupying a similar geographic range in North America ( Asquith, 1990).

SYNTYPE SERIES: Uhler (1894) in the original description of A. princeps stated that it was found in ‘‘northern California, Oregon and Washington State’’. Specimens from Uhler’s collection in the USNM and AMNH are from various western localities: ‘‘Cal.’’ (California), ‘‘W.T.’’ (Western Territories), ‘‘Klikitat V., W.T.’’ (Klickitat Valley, Western Territories), ‘‘Yakima City, W.T.’’ (Western Territories), and ‘‘Yakima R., W.T.’’ (Yakima River, Western Territories). All of those specimens fit the color description made by Uhler. Of those, there is one in the USNM labeled as ‘‘ Hadronema princeps ’’ with handwriting similar to that of Uhler and with a label ‘‘COTYPE (by Uhler) ‘ princeps ’’’ affixed by Knight. This specimen is almost complete, and it has been repinned on a point, probably also by Knight. Because this male is in good condition, and it fits Uhler’s description and distributional data, it is designated as the lectotype for A. princeps .

LECTOTYPE MALE (here designated): USA: Washington: Yakima Co.: Yakima R.[iver], W.[estern] T.[erritories], [46.49984 ° N 120.4503 ° W], 04 Jul 1882 – 05 Jul 1882, Nelson’s, P.R. Uhler collection, ‘‘ Hadronema princeps Uhler W.T. ’’, Cotype (by Uhler) ‘‘ princeps ’’, Lectotype Hadronema princeps D. Forero det. (red label), 13 (AMNH_PBI 00106738) (USNM).

HOLOTYPES OF SYNONYMS: USA: California: San Diego Co.: San Diego County, 32.71528 ° N 117.15639 ° W, 08 Jun 1913, E.P. Van Duzee, 13 (AMNH_PBI 00077855) ( Aoplonema uhleri ) (CAS). Oregon: Malheur Co.: Vale, 43.9821 ° N 117.2382 ° W, 684 m, 16 Jun 1926, R.W. Haegele, greasewood, 64, Holotype (by H.H. Knight) ‘‘ Hadronema uniformis ’’, H.H. Knight coll. 1976, 13 (AMNH_PBI 00070388) ( Aoplonema uniforme ) (USNM).

PARALECTOTYPES: USA: California: Unknown Co.: Unknown locality, 2♀ ( AMNH _ PBI 00107137 , AMNH _ PBI 00107138 ) ( USNM) . Washington: Klickitat Co.: Kli [c]kitat V.[alley], W.[estern] T.[erritories], 45.81706 ° N 121.1525 ° W, 146 m, 11 Jul 1882, unknown collector, 13 ( AMNH _ PBI 00102485 ) ( AMNH). 23 ( AMNH _ PBI 00107122 , AMNH _ PBI 00107123 ) ( USNM) ; 10 Jul 1882, Thorpe , 13 ( AMNH _ PBI 00102486 ), 1♀ ( AMNH _ PBI 00102489 ) ( AMNH). 1♀ ( AMNH _ PBI 00107136 ) ( USNM) . Yakima Co.: Yakima City , W.T. [Western Territories], 46.55734 ° N 120.475 ° W, 02 Jul 1882 – 04 Jul 1882, unknown collector, 1♀ ( AMNH _ PBI 00107135 ) ( USNM). Yakima R.[iver], W.[estern] T.[erritories], 46.49984 ° N 120.4503 ° W, 04 Jul 1882 – 05 Jul 1882, Nelson, 1♀ ( AMNH _ PBI 00102490 ) ( AMNH) .

PARATYPES (OF SYNONYMS): USA: California: San Diego Co.: San Diego County, 32.71528 ° N 117.15639 ° W, 23 May 1913, E.P. Van Duzee, Paratype of A. uhleri , 23, 2♀ (13, 1♀ on each pin) (AMNH_PBI 00077863, AMNH_PBI 00077864) (CAS); 24 May 1913, E.P. Van Duzee, Paratype of A. uhleri , 23 (AMNH_PBI 00077862, AMNH_PBI 00077865) (CAS); 19 Jun 1913, E.P. Van Duzee, Paratype of A. uhleri , 13 (AMNH_PBI 00077866) (CAS); 21 Jun 1914, E.P. Van Duzee, Paratype of A. uhleri , 13 (AMNH_PBI 00077861) (CAS). Idaho: Canyon Co.: Caldwell, 43.66294 ° N 116.68735 ° W, 15 Jun 1926, R.W. Haegele, Paratype of A. uniforme , 13 (AMNH_PBI 00104807) (CNC). Oregon: Malheur Co.: Vale, 43.9821 ° N 117.2382 ° W, 684 m, 16 Jun 1926, R.W. Haegele, Paratype of A. uniforme , Sarcobatus sp. (Sarcobataceae) , 1♀ (AMNH_ PBI 00106621) (USNM).

OTHER SPECIMENS EXAMINED: CANADA: Alberta: Blairmore, 49.6 ° N 114.43333 ° W, 14 Aug 1930, J.H. Pepper, 1♀ (AMNH_PBI 00104946) (CNC). Cypress Hills Provincial Park, Murray Hill Road, 49.63 ° N 110.2 ° W, 1433 m, 18 Jul 1990, M.D. Schwartz, 1♀ (AMNH_PBI 00104923) (CNC). Cypress Hills Provincial Park, Top Road, 9.6 km E of Rt 41 jct, 49.63 ° N 110.2 ° W, 1433 m, 15 Jul 1990, M.D. Schwartz, 13 (AMNH_PBI 00104921), 1♀ (AMNH_PBI 00104922) (CNC). Elkwater Park, Cypress Hills Provincial Park, 49.63 ° N 110.2 ° W, 13 Jul 1952, A.R. Brooks, 13 (AMNH_PBI 00104904) (CNC); 29 Jul 1952, L.A. Konotopetz, 1♀ (AMNH_PBI 00104908) (CNC); 16 Jul 1952, A.R. Brooks, 23 (AMNH_PBI 00104903, AMNH_PBI 00104905) (CNC); 18 Jul 1952, A.R. Brooks, 1♀ (AMNH_PBI 00104907) (CNC). Kananaskis Hwy, 50.91555 ° N 115.14166 ° W, 25 Jul 1973, L.A. Kelton, 13 (AMNH_PBI 00104924), 1♀ (AMNH_PBI 00104943) (CNC). Lethbridge, 49.7 ° N 112.83333 ° W, 18 Jul 1930, J.H. Pepper, 1♀ (AMNH_PBI 00104945) (CNC); 15 Jul 1928, J. McDunnough, 1♀ (AMNH_PBI 00104944) (CNC). Lundbreck, 49.58 ° N 114.17 ° W, 07 Jul 1970, L.A. Kelton, 2♀ (AMNH_PBI 00104947, AMNH_PBI 00104948) (CNC). Manyberries, 49.4 ° N 110.7 ° W, 08 Jul 1952, L.A. Konotopetz, Juniperus sp. (Cupressaceae) , 63 (AMNH_ PBI 00104909–AMNH_PBI 00104914), 6♀ (AMNH_PBI 00104915–AMNH_PBI 00104920) (CNC). Medicine Hat, 50.03333 ° N 110.68333 ° W, 15 Jun 1930, J.H. Pepper, 1♀ (AMNH_PBI 00104949) (CNC). Waterton Lakes National Park, 49.05 ° N 113.9 ° W, 22 Jul 1923, J. McDunnough, 1♀ (AMNH_PBI 00104941) (CNC). Waterton Park, 49.05 ° N 113.91666 ° W, 26 Jul 1972, L.A. Kelton, Pinus contorta (Pinaceae) , 43 (AMNH_PBI 00104930–AMNH_PBI 00104933) Juniperus horizontalis (Cupressaceae) , 33 (AMNH_PBI 00104934–AMNH_ PBI 00104936), 53 (AMNH_PBI 00104925– AMNH_PBI 00104929), 2♀ (AMNH_PBI 00104938, AMNH_PBI 00104939) (CNC); 14 Jul 1923, E.H. Strickland, 13 (AMNH_ PBI 00104937) (CNC); 18 Jul 1952, A.R. Brooks, 1♀ (AMNH_PBI 00104940) (CNC). British Columbia : Caribou Co.: Soda Creek, 52.33333 ° N 122.28333 ° W, 22 Jul 1947, G.J. Spencer, 23 (AMNH_PBI 00104830, AMNH_ PBI 00104831) (CNC); 12 Jul 1950, G.J. Spencer, 23 (AMNH_PBI 00104825, AMNH_PBI 00104826), 6♀ (AMNH_PBI 00104833–AMNH_ PBI 00104838) (CNC); 04 Jul 1950, G.J. Spencer, 13 (AMNH_PBI 00104827) (CNC); 27 Jul 1949, G.J. Spencer, 23 (AMNH_PBI 00104828, AMNH_PBI 00104829) (CNC); 08 Jul 1949, G.J. Spencer, 13 (AMNH_PBI 00104832) (CNC). 10 mi S of Merritt, 49.97209 ° N 120.78333 ° W, 19 Jul 1959, L.A. Kelton, 1♀ (AMNH_PBI 00104898) (CNC). 10 mi W of Princeton, 49.45 ° N 120.72237 ° W, 31 Jul 1957, N.H. Anderson, 13 (AMNH_PBI 00104891) (CNC). Elko, 49.3 ° N 115.11666 ° W, 08 Jul 1970, L.A. Kelton, 1♀ (AMNH_PBI 00104899) (CNC). Midday Val[ley]., Merritt, 49.98333 ° N 120.93333 ° W, 736 m, Jul 1924, K.F. Auden, 4♀ (AMNH_PBI 00104894–AMNH_PBI 00104897) (CNC). Naramata, 49.59502 ° N 119.59556 ° W, 28 May 1958, H. & A. Howden, 1♀ (AMNH_PBI 00104892) (CNC); 16 Jul 1950, B.P. Beirne, 1♀ (AMNH_PBI 00104893) (CNC). Oliver, 49.18333 ° N 119.55 ° W, 19 Jul 1970, L.A. Kelton, 1♀ (AMNH_PBI 00104866) (CNC); 02 Jul 1974, L.A. Kelton, Ceanothus sp. (Rhamnaceae) , 13 (AMNH_ PBI 00104839), 3♀ (AMNH_PBI 00104840– AMNH_PBI 00104842) (CNC); 02 Jul 1975, L.A. Kelton, Lupinus sp. (Fabaceae) , 1♀ (AMNH_PBI 00104843) (CNC); 29 Jun 1959, L.A. Kelton, 1♀ (AMNH_PBI 00104875) (CNC). Oliver, 7 mi E of Indian Reservation, 49.48904 ° N 119.42862 ° W, 08 Jul 1959, L.A. Kelton, Salvia sp. (Lamiaceae) , 73 (AMNH_ PBI 00104867–AMNH_PBI 00104873), 7♀ (AMNH_PBI 00104876–AMNH_PBI 00104882) (CNC). Oliver, McIntyre Creek, 49.18333 ° N 119.55 ° W, 04 Jul 1959, L.A. Kelton, 33 (AMNH_PBI 00104820–AMNH_PBI 00104822), 2♀ (AMNH_PBI 00104823, AMNH_PBI 00104824) (CNC). Oliver, McKinney Road, 49.18333 ° N 119.55 ° W, 08 Jul 1959, L.A. Kelton, 1♀ (AMNH_PBI 00104900) (CNC). Oliver, Sawmill Lake, 49.18333 ° N 119.55 ° W, 30 Jun 1959, L.A. Kelton, Lupinus sp. (Fabaceae) , 23 (AMNH_PBI 00104889, AMNH_PBI 00104890) (CNC). Oliver, Vaseaux Lake, 49.18333 ° N 119.55 ° W, 26 Jun 1959, L.A. Kelton, 13 (AMNH_PBI 00104888) (CNC); 10 Jul 1959, L.A. Kelton, Sarcobatus sp. (Sarcobataceae) , 13 (AMNH_PBI 00104887) (CNC). Oliver, White Lake, 49.18333 ° N 119.55 ° W, 29 Jun 1959, L.A. Kelton, Lupinus sp. (Fabaceae) , 13 (AMNH_PBI 00104852), 1♀ (AMNH_ PBI 00104853) (CNC). Osoyoos, Anarchist Mountain, 49.03333 ° N 119.33333 ° W, 20 Jul 1959, L.A. Kelton, 1♀ (AMNH_PBI 00104883) (CNC); 13 Jul 1970, L.A. Kelton, 83 (AMNH_ PBI 00104854–AMNH_PBI 00104861), 4♀ (AMNH_PBI 00104863–AMNH_PBI 00104865, AMNH_PBI 00104874) (CNC); 06 Jul 1959, R. Madge, 13 (AMNH_PBI 00104884) (CNC). Summerland, 49.56646 ° N 119.63951 ° W, 400 m, 06 Jul 1975, L.A. Kelton, 1♀ (AMNH_PBI 00104851) (CNC); 10 Jul 1975, L.A. Kelton, light trap, 13 (AMNH_PBI 00104862) (CNC); 12 Jul 1975, L.A. Kelton, 2♀ (AMNH_PBI 00104846, AMNH_PBI 00104847) (CNC), Ceanothus sp. (Rhamnaceae) , 3♀ (AMNH_PBI 00104848– AMNH_PBI 00104850); 02 Jul 1974 – 11 Jul 1974, L.A. Kelton, Ceanothus sp. (Rhamnaceae) , 23 (AMNH_PBI 00104844, AMNH_ PBI 00104845) (CNC). Texas creek, Lillooet, 50.56666 ° N 121.8 ° W, 15 Jun 1926, J. McDunnough, Chrysothamnus sp. (Asteraceae) , 113 (AMNH_PBI 00104808–AMNH_PBI 00104818), 1♀ (AMNH_PBI 00104819) (CNC). Vernon, 50.26666 ° N 119.26666 ° W, 02 Jul 1920, M.H. Ruhmann, 1♀ (AMNH_PBI 00107184) (USNM); 21 Jul 1923, D.G. Gillespie, 1♀ (AMNH_PBI 00104901) (CNC). Williams Lake, 52.11666 ° N 122.15 ° W, 12 Jul 1938, J.K. Jacob, 13 (AMNH_ PBI 00104885), 1♀ (AMNH_PBI 00104886) (CNC). Saskatchewan: Val Marie, 49.23333 ° N 107.73333 ° W, 08 Aug 1955, A.R. Brooks, 1♀ (AMNH_PBI 00104942) (CNC). MEXICO: Baja California: 38 km E Rt 1 to Parque San Pedro Martir, 30.96 ° N 115.82 ° W, 400 m, 24 Apr 1985, R.T. Schuh and B.M. Massie, 103 (AMNH_PBI 00102212, AMNH_PBI 00102215, AMNH_PBI 00102217–AMNH_ PBI 00102224), 11♀ (AMNH_PBI 00102225– AMNH_PBI 00102228, AMNH_PBI 00102232– AMNH_PBI 00102238), Viguiera laciniata A. Gray (Asteraceae) , 2♀ (AMNH_PBI 00102229, AMNH_PBI 00102230), Lotus scoparius (Nott.) Ottley (Fabaceae) , 43 (AMNH_PBI 00102214, AMNH_PBI 00102216, AMNH_PBI 00102765, AMNH_PBI 00102784), 1♀ (AMNH_PBI 00102231), Sphaeralcea emoryi Torr. (Malvaceae) , 23 (AMNH_PBI 00102213, AMNH_ PBI 00102239), 7♀ (AMNH_PBI 00102240– AMNH_PBI 00102246) (AMNH). USA: Arizona: Apache Co. : Eagar, ApacheNational Forest, 34.11111 ° N 109.29083 ° W, 2286 m, 12 Jul 1968, L. A. Kelton, 13 (AMNH_PBI 00104957) (CNC). Coconino Co.: Williams, 35.24944 ° N 112.19028 ° W, H.S. Barber, 13 (AMNH_PBI 00107237), 1♀ (AMNH_PBI 00107239) (USNM); collector unknown, 13 (AMNH_PBI 00107238) (USNM). California: El Dorado Co.: Phillips, 38.81777 ° N 120.0775 ° W, 24 Aug 1968, W.F. Chamber- lain, 1♀ (AMNH_PBI 00106289) (TAMU). Fresno Co.: San Benito, Big Panoche Creek, 36.74841 ° N 120.51356 ° W, 21 Apr 1967, J. Powell, 53 (AMNH_PBI 00079863– AMNH_PBI 00079867), 7♀ (AMNH_PBI 00079868–AMNH_PBI 00079874) (UCB). Inyo Co. : E side of California Rt 141, 39.00944 ° N 120.76972 ° W, 1524 m, 31 May 1981, J.T. Polhemus, 1♀ (AMNH_PBI 00065089) (JTP). Kern Co.: Near Walker Pass, 35.6625 ° N 118.02583 ° W, 1524 m, 30 May 1981, J.T. Polhemus, 2♀ (AMNH_PBI 00065087, AMNH_PBI 00065088) (JTP). Los Angeles Co.: Los Angeles County, 34.05222 ° N 118.24278 ° W, Coquillett Collection, 13 (AMNH_PBI 00106615) (USNM). Mariposa Co. : Yosemite National Park, 37.85 ° N 119.56667 ° W, 1201 m, 29 May 1931, collector unknown, 13 (AMNH_PBI 00107170) (USNM). Modoc Co.: Upper Alkali Lake, Warner Mountains, 41.25222 ° N 120.60389 ° W, 02 Aug 1922, C.L. Fox, 1♀ (AMNH_PBI 00106619) (USNM). Mono Co.: Mono Lake, N shore, 38.07365 ° N 118.99159 ° W, 09 Aug 1997, G.R. Ballmer, Artemisia tridentata (Asteraceae) , 1♀ (AMNH_PBI 00083531) (UCR). Monterey Co.: Pleyto, 35.86028 ° N 120.9925 ° W, 21 May 1920, E.P. Van Duzee, 13 (AMNH_PBI 00077869) (CAS). Riverside Co.: 5.6 mi S of Sage on Route 3, site 2, T7S R1E S32, 33.51666 ° N 116.9 ° W, 04 Jun 1976, collector unknown, 13 (AMNH_PBI 00083125) (UCR). Pauba Valley, T8S R1W S19, 33.5015 ° N 117.0307 ° W, 366 m, 12 May 1987, R.D. Goeden and D.W. Ricker, Encelia virginensis (Asteraceae) , 23 (AMNH_PBI 00083118, AMNH_PBI 00083119) (UCR). Encelia virginensis (Asteraceae) , 23 (AMNH_ PBI 00107228, AMNH_PBI 00107229) (USNM). Riverside, 33.95333 ° N 117.39528 ° W, 22 May 1933, Timberlake, Astragalus purshii (Fabaceae) , 13 (AMNH_PBI 00083124) (UCR). Temecula Canyon, Santa Margarita River , 33.41417 ° N 117.24389 ° W, 229 m, 18 May 1968, L.O. Tejada, Lotus sp. (Fabaceae) , 13 (AMNH_PBI 00083081) (UCR). San Bernardino Co.: Colton, 34.07389 ° N 117.31278 ° W, 26 May 1917 – 28 May 1917, E.P. Van Duzee, 33 (AMNH_PBI 00077870–AMNH_PBI 00077872) (CAS). Joshua Tree National Monument, Covington Flat, 34.0089 ° N 116.3125 ° W, 17 Jun 1977, K.W. Cooper, 23 (AMNH_PBI 00083126, AMNH_PBI 00083127) (UCR). San Diego Co.: Guatay, 32.84893 ° N 116.5572 ° W, 19 Jul 1941, L.H. Banker, 13 (AMNH_PBI 00075266) (KU). Mission Dam, 32.84199 ° N 117.04058 ° W, 04 Jun 1965, J. Powell, 13 (AMNH_PBI 00079882) (UCB). Pine Hills, 33.01616 ° N 116.63446 ° W, 1108 m, 22 Jun 1950, Timberlake, Symphoricarpos albus (Caprifoliaceae) , 13 (AMNH_ PBI 00083122) (UCR). Siskiyou Co.: 5 mi S of Merrill, Oregon, near Tule Lake, 41.96785 ° N 121.56512 ° W, 1237 m, 26 Jun 1979, M.D. Schwartz, Sarcobatus vermiculatus (Sarcobataceae) , 83 (AMNH_PBI 00102251–AMNH_PBI 00102258), 22♀ (AMNH_ PBI 00102263–AMNH_PBI 00102283, AMNH_ PBI 00102755) (AMNH); 26 Jun 1979, G.M. Stonedahl, Sarcobatus vermiculatus (Sarcobataceae) , 10♀ (AMNH_PBI 00102285– AMNH_PBI 00102294) (AMNH). McCloud, 41.25583 ° N 122.13833 ° W, 1090 m, 23 Jul 1918, E.P. Van Duzee, 13 (AMNH_PBI 00107171) (USNM). Siskiyou County, 41.58333 ° N 122.51667 ° W, collector unknown, 1♀ (AMNH_PBI 00107151) (USNM). East side of Lower Klamath Lake, 41.99145 ° N 121.71243 ° W, 1270 m, 26 Jun 1979, R.T. and Joe Schuh, Sarcobatus vermiculatus (Sarcobataceae) , 23 (AMNH_PBI 00102259, AMNH_ PBI 00102779), 7♀ (AMNH_PBI 00102260– AMNH_PBI 00102262, AMNH_PBI 00102590– AMNH_PBI 00102593) (AMNH). Tulare Co.: 15 mi N of Kernville, 35.96844 ° N 118.47937 ° W, 10 May 1997, W.F. Chamberlain, 13 (AMNH_ PBI 00105586) (TAMU). 4.5 mi S of Three Rivers, 36.37363 ° N 118.90361 ° W, 29 Apr 1947, Timberlake, Frangula californica (Rhamnaceae) , 13 (AMNH_PBI 00083120) (UCR). California Hot Springs, 35.88028 ° N 118.67278 ° W, Jun 1953, Timberlake, Eriodictyon californicum (Hydrophyllaceae) , 13 (AMNH_PBI 00083121) (UCR). Sequoia National Park, Ash Mountain Research Center, site 2, 36.49194 ° N 118.83056 ° W, 08 May 1987 – 18 May 1987, S. Haultain-Tweed, in cantharidin trap, 23 (AMNH_PBI 00133862, AMNH_PBI 00133863), 2♀ (AMNH_PBI 00133864, AMNH_PBI 00133865) (USNM). Unknown Co.: Unknown locality, 08 May 1950, Timberlake, Eriogonum fasciculatum (Polygonaceae) , 13 (AMNH_PBI 00083123) (UCR). Colorado: Arapahoe Co.: Cherry Creek State Park, 39.63118 ° N 104.85283 ° W, 07 Jun 1977, D.A. and J.T. Polhemus, 13 (AMNH_PBI 00065142) (JTP). Boulder Co.: 6 mi W of Boulder, 40.015 ° N 105.38325 ° W, 12 Aug 1973, J.C. Schaffner, 1♀ (AMNH_ PBI 00106293) (TAMU). Boulder, 40.015 ° N 105.27 ° W, 1676 m, 21 Jul 1903, E.P. Van Duzee, 13 (AMNH_PBI 00077887) (CAS); 29 Jun 1932, C.H. Hicks, 4♀ (AMNH_PBI 00107214–AMNH_PBI 00107215, AMNH_ PBI 00107217, AMNH_PBI 00107218) (USNM); 18 Jul 1932, C.H. Hicks, 1♀ (AMNH_PBI 00107201) (USNM); 17 Jun 1932, C.H. Hicks, 43 (AMNH_PBI 00107209– AMNH_PBI 00107212) (USNM); 16 Jun 1932, C.H. Hicks, 2♀ (AMNH_PBI 00107213, AMNH_PBI 00107216) (USNM). Nederland, Science Lodge, 39.96139 ° N 105.51028 ° W, 2743 m, 29 Jul 1961, J.R. Stainer, 1♀ (AMNH_PBI 00104956) (CNC). Nederland, Science Lodge, 39.96139 ° N 105.51028 ° W, 2896 m, 06 Jul 1961, J.R. Stainer, 13 (AMNH_PBI 00104950) (CNC). Valmont Butte, 40.03081 ° N 105.2116 ° W, 1615 m, 30 Jul 1961, J.R. Stainer, 1♀ (AMNH_PBI 00104955) (CNC). Clear Creek Co.: Timberline, II, Mt. Evans, 39.58859 ° N 105.64333 ° W, 183 m, 21 Jul 1961, J.R. Stainer, 13 (AMNH_PBI 00104951) (CNC). Dolores Co.: 29 mi SW of Norwood, 37.8341 ° N 108.66702 ° W, 07 Jul 1980, J.T. and D.A. Polhemus, 1♀ (AMNH_PBI 00065109) (JTP). Douglas Co.: Chatfield State Park, 39.53666 ° N 105.06888 ° W, 03 Jun 1992, J.T. Polhemus, 33 (AMNH_PBI 00065104–AMNH_PBI 00065106), 1♀ (AMNH_PBI 00065107) (JTP); 10 Jun 1977, D.A. and J.T. Polhemus, 13 (AMNH_PBI 00065132) (JTP). Head of Highline Canal, 39.56168 ° N 104.99692 ° W, 02 Jul 1979, J.T. Polhemus, 13 (AMNH_PBI 00065131) (JTP); 16 Jun 1978, J.T. Polhemus, 23 (AMNH_PBI 00065117, AMNH_PBI 00065141), 5♀ (AMNH_PBI 00065108, AMNH_PBI 00065134–AMNH_PBI 00065137) (JTP); 20 Jun 1978, J.T. Polhemus, 13 (AMNH_PBI 00065092), 1♀ (AMNH_PBI00065138) (JTP); 02 Jul 1979, J.T. Polhemus, 33 (AMNH_PBI 00065093–AMNH_ PBI 00065095) (JTP); 03 Jul 1979, J.T. Polhemus, 23 (AMNH_PBI 00065101, AMNH_PBI 00065140), 1♀ (AMNH_PBI 00065096) (JTP). Perry Park, 39.25667 ° N 104.99194 ° W, 13 Jul 1977, D.A. and J.T. Polhemus, 33 (AMNH_PBI 00065110–AMNH_PBI 00065112), 2♀ (AMNH_ PBI 00065113, AMNH_PBI 00065114) (JTP), 1♀ (AMNH_PBI 00107200) (USNM). Roxborough Park Road near Chatfield State Park, 39.47389 ° N 105.08472 ° W, 1707 m, 06 Jul 1979, J.T. Polhemus, 13 (AMNH_PBI 00065143) (JTP). Waterton, 39.49361 ° N 105.08806 ° W, 10 Jun 1982, D.A. Polhemus, Rhus trilobata (Anacardiaceae) , 13 (AMNH_PBI 00065128), 1♀ (AMNH_PBI 00065130) (JTP); 07 Jul 1983, D.A. Polhemus, 13 (AMNH_PBI 00065126) (JTP); 17 Jun 1982, D.A. Polhemus, 13 (AMNH_PBI 00065127) (JTP); 22 Jun 1981, J.T. Polhemus, 13 (AMNH_PBI 00065079) (JTP); 01 Jul 1986, D.A. Polhemus, 13 (AMNH_PBI 00065080), 1♀ (AMNH_PBI 00065086) (JTP); 13 Jun 1986, D.A. Polhemus, 13 (AMNH_PBI 00065081) (JTP); 16 Jul 1981, J.T. Polhemus, 1♀ (AMNH_PBI 00065082) (JTP); 13 Jun 1985, D.A. Polhemus, Ribes cereum (Grossulariaceae) , 13 (AMNH_PBI 00065119) (JTP); 24 Jun 1982, D.A. Polhemus, 13 (AMNH_PBI 00065129) (JTP). Waterton, Head of Hiline, 39.49361 ° N 105.08806 ° W, 18 Jul 1979, J.T. Polhemus, 13 (AMNH_PBI 00065097) (JTP); 17 Jun 1980, J.T. Polhemus, 13 (AMNH_PBI 00065102) (JTP); 13 Jun 1980, J.T. Polhemus, 13 (AMNH_PBI 00065098) (JTP); 20 Jun 1980, J.T. Polhemus, 13 (AMNH_PBI 00065099), 1♀ (AMNH_PBI 00065100) (JTP). Near Waterton, Roxborough Road, 39.49361 ° N 105.08806 ° W, 1707 m, 11 Jun 1981, D.A. Polhemus, 1♀ (AMNH_PBI 00065085) (JTP). Eagle Co.: 3.9 mi S of Minturn, White River National Forest, 39.52986 ° N 106.43028 ° W, 2859 m, 19 Jul 1990, G. Zolnerowich, 13 (AMNH_PBI 00106291) (TAMU). Grouse Creek trail, N of Minturn, 39.59554 ° N 106.43336 ° W, 2390 m, 11 Aug 1986, J.T. and D.A. Polhemus, 1♀ (AMNH_ PBI 00065116) (JTP). Vail, 39.64028 ° N 106.37361 ° W, 26 Jun 1977, J.T. Polhemus, 1♀ (AMNH_PBI 00065139) (JTP). Vail, 39.64 ° N 106.37416 ° W, 23 Jun 1986, J.T. Polhemus, 1♀ (AMNH_PBI 00065115) (JTP). Garfield Co.: Grizzly Creek N. (Colo. 2193), 39.7057 ° N 107.27825 ° W, 19 Jul 1896, C.F. Baker, Artemisia tridentata (Asteraceae) , 1♀ (AMNH_ PBI 00107147) (USNM). Gilpin Co.: Rollinsville, Roosevelt National Forest, 39.91722 ° N 105.50056 ° W, 02 Sep 1972, L.A. Kelton, 3♀ (AMNH_PBI 00104952–AMNH_PBI 00104954) (CNC). Jefferson Co.: Deer Creek Canyon, 39.85288 ° N 105.33944 ° W, 1981 m, 11 Jul 1986, R.T. Schuh and J.T. Polhemus, 183 (AMNH_PBI 00102547–AMNH_PBI 00102563, AMNH_PBI 00102782), 3♀ (AMNH_PBI 00102564–AMNH_PBI 00102566) (AMNH). Golden, 39.75556 ° N 105.22056 ° W, 1829 m, 25 Jul 1909, W.J. Gerhard, 13(AMNH_PBI00107193) (USNM); 24 Jul 1909, W.J. Gerhard, 1♀ (AMNH_PBI 00107199) (USNM). Indian Hills, 39.61667 ° N 105.23667 ° W, 2134 m, 11 Jul 1986, R.T. Schuh and J.T. Polhemus, Astragalus sp. (Fabaceae) , 33 (AMNH_PBI 00101307– AMNH_PBI 00101309), 10♀ (AMNH_PBI 00101314–AMNH_PBI 00101322, AMNH_ PBI 00102754) (AMNH). Lookout Mountain, 39.7324 ° N 105.238 ° W, 2134 m, 21 Jul 1916, A.C. Burrill, Penstemon sp. (Scrophulariaceae) , 13 (AMNH_PBI 00107192) (USNM). North Turkey Creek Park near Tenders, 39.59468 ° N 105.22014 ° W, 1890 m, 16 Jul 1983, R.T. Schuh, D.A. and J.T. Polhemus, 33 (AMNH_PBI 00101306, AMNH_PBI 00102778, AMNH_PBI 00102783), 1♀ (AMNH_PBI 00101313) (AMNH). Red Rocks Park, 39.66972 ° N 105.20278 ° W, 07 Jul 1983, D.A. Polhemus, 23 (AMNH_PBI 00065120, AMNH_PBI 00065121) (JTP); 23 Jun 1982, D.A. Polhemus, 43 (AMNH_PBI 00065122–AMNH_PBI 00065125) (JTP); 21 Jun 1988, D.A. Polhemus, 1♀ (AMNH_PBI 00065083) (JTP); 29 Jun 1988, J.T. Polhemus, 1♀ (AMNH_PBI 00065084) (JTP). Red Rocks Park near Morrison, 39.65361 ° N 105.19056 ° W, 1707 m, 15 Jul 1983, R.T. Schuh and D.A. Polhemus, 113 (AMNH_ PBI 00101295–AMNH_PBI 00101305), 3♀ (AMNH_PBI 00101310–AMNH_PBI 00101312) (AMNH). Larimer Co.: 40 mi W Fort Collins, Bennett Crk. Picnic Ground, Pingree Pk. Rd., 40.58 ° N 105.847 ° W, 2256 m, 14 Jul 1986, R.T. Schuh and J.T. Polhemus, 13 (AMNH_ PBI 00102640) (AMNH). Forrester’s (Colo. 2020, Colo. 1581), 40.93952 ° N 105.98546 ° W, 19 Jul 1895, C.F. Baker, 23 (AMNH_PBI 00125487, AMNH_PBI 00125488), 1♀ (AMNH_ PBI 00125489) (CUIC). 13 (AMNH_PBI 00107194), 1♀ (AMNH_PBI 00107148) (USNM). Forrester’s Ranch (Colo. 2013), 40.93952 ° N 105.98546 ° W, 03 Aug 1896, C.F. Baker, 1♀ (AMNH_PBI 00107139) (USNM). Fort Collins, 40.58528 ° N 105.08389 ° W, 13 Jun 1898, collector unknown, 13 (AMNH_PBI 00075272) (KU); 07 Jun 1899, collector unknown, 13 (AMNH_PBI 00075284) (KU); 09 Jul 1902, collector unknown, 13 (AMNH_PBI 00107195) (USNM); 17 Jul 1900, E.P. Van Duzee, 13 (AMNH_PBI 00077886), 1♀ (AMNH_PBI 00077888) (CAS). Prairie Divide, 40.85553 ° N 105.4922 ° W, 06 Aug 1965, collector unknown, 13 (AMNH_PBI 00106292) (TAMU). Rocky Mountain National Park, Moraine Valley, 40.33333 ° N 105.70833 ° W, 2438 m, 08 Jul 1966, J.A. Slater, 1♀ (AMNH_PBI 00102568) (AMNH). Las Animas Co.: 1 mi N of Stonewall on Purgatoire Campground Road, 37.16667 ° N 105.01667 ° W, 2560 m, 18 Aug 1986, R.T. Schuh, 1♀ (AMNH_PBI 00102567) (AMNH). Mineral Co.: Creede, 37.84917 ° N 106.92583 ° W, 04 Jul 1977, J.T. Polhemus, 13 (AMNH_PBI 00065133) (JTP). Montezuma Co. : Cortez, 37.34889 ° N 108.58528 ° W, 19 Jul 1968, L.A. Kelton, Salicornia sp. (Chenopodiaceae) , 1♀ (AMNH_PBI 00105193) (CNC). Dolores, 37.47389 ° N 108.50389 ° W, 15 Aug 1925, H.H. Knight, 1♀ (AMNH_PBI 00107180) (USNM). Montrose Co.: 25 mi E of Nucla on Uncompahgre Plateau, 38.4255 ° N 108.4222 ° W, 2804 m, 14 Aug 1987, J.T. and D.A. Polhemus, 13 (AMNH_PBI 00065078) (JTP). 27 mi E of Nucla, on Uncompahgre Plateau, 38.26944 ° N 108.05005 ° W, 14 Aug 1987, J.T. and D.A. Polhemus, 1♀ (AMNH_PBI 00065077) (JTP). Routt Co.: 8 mi S of Steamboat Springs (Colo. 2030), 40.36906 ° N 106.83111 ° W, 21 Jul 1896, Liebeck, 23 (AMNH_PBI 00107142, AMNH_PBI 00107143), 1♀ (AMNH_PBI 00107203) (USNM). Steamboat Springs, 40.485 ° N 106.83111 ° W, 2103 m, 01 Jul 1944, collector unknown, 1♀ (AMNH_PBI 00075261) (KU); 29 Jun 1931, E.D. Ball, 13 (AMNH_PBI 00107198) (USNM). Unknown Co.: Colo. 1685, collector unknown, 1♀ (AMNH_PBI 00125490) (CUIC). Colo. 1693, O. Heidemann, 13 (AMNH_PBI 00125491) (CUIC); collector unknown, 13 (AMNH_PBI 00077885) (CAS). Colo. 1732, O. Heidemann, 13 (AMNH_PBI 00125486) (CUIC). Colo. 2012, C.F. Baker, 13 (AMNH_PBI 00107191) (USNM). Colo. 2214, collectorunknown, 1♀ (AMNH_PBI 00077889) (CAS). Colo. 795, collector unknown, 13 (AMNH_ PBI 00107197) (USNM). Dtch Geo, 11 Aug 1899, collector unknown, 1♀ (AMNH_PBI 00075273) (KU). Idaho: Adams Co.: Council, 44.72988 ° N 116.4381 ° W, 892 m, 25 Jun 1926, R.W. Haegele, 13 (AMNH_PBI 00107154) (USNM). Mesa, 44.62877 ° N 116.4507 ° W, 991 m, 13 Jul 1936, W.E. Schull, 13 (AMNH_ PBI 00107155) (USNM). Bannock Co.: 3 mi E of McCammon, 42.65056 ° N 112.13325 ° W, 29 Jun 1966, W. Gagné and J. Haddock, 1♀ (AMNH_PBI 00079875) (UCB). Bear Lake Co.: Georgetown, 42.48214 ° N 111.3707 ° W, 06 Jul 1951, G.F. Knowlton, 1♀ (AMNH_ PBI 00107150) (USNM). Paris, F 4741, 42.23333 ° N 111.41666 ° W, 1829 m, 08 Jul 1920, collector unknown, 13 (AMNH_PBI 00102462) (AMNH). Bonneville Co.: Grays Lake National Wildlife Refuge, road to lookout, 43.05144 ° N 111.38204 ° W, 30 Jul 1981, M.D. Schwartz, 23 (AMNH_PBI 00102491, AMNH_PBI 00102492), 2♀ (AMNH_ PBI 00102493, AMNH_PBI 00102494) (AMNH). Butte Co.: Craters of the Moon National Monument, 43.41667 ° N 113.51667 ° W, 28 Jul 1965, D.S. Horning, 1♀ (AMNH_PBI 00107181) (USNM). Clearwater Co.: Weippe, 46.37601 ° N 115.9381 ° W, 919 m, 05 Jul 1935, W.E. Schull, 13 (AMNH_PBI 00107159), alfalfa, 13 (AMNH_PBI 00107158) (USNM). Franklin Co.: Dayton, 42.11306 ° N 111.99278 ° W, 1447 m, 29 Jul 1937, R.E. Miller, 1♀ (AMNH_PBI 00107186) (USNM). Williams Canyon, mp 20 on Rt 36, T12S R42 S30, 42.35694 ° N 111.69583 ° W, 2438 m, 19 Jul 1981, M.D. Schwartz, Rudbeckia occidentalis (Asteraceae) , 13 (AMNH_PBI 00102514) (AMNH). Fremont Co.: 14 mi N of Ashton, 44.27444 ° N 111.4475 ° W, 22 Jul 1991, R.S. Peigler, 13 (AMNH_PBI 00105611), 1♀ (AMNH_PBI 00105612) (TAMU). Henrys Lake, 44.59722 ° N 111.3525 ° W, 2134 m, 12 Jul 1936, R.E. Miller, 13 (AMNH_PBI 00107156) (USNM). Gem Co.: Emmett, 43.87361 ° N 116.49833 ° W, 716 m, 23 Jun 1923, R.W. Haegele, Sarcobatus sp. (Sarcobataceae) , 13 (AMNH_PBI 00106618) (USNM). Idaho Co.: Lolo Pass, at junction of Route 12 & FS Road 595 (to Granite Pass), Clearwater National Forest, 46.63514 ° N 114.57952 ° W, 30 Jul 1994, M.D. Schwartz, ( Asteraceae ), 1♀ (AMNH_PBI 00104902) (CNC). Latah Co.: Moscow, 46.7325 ° N 116.99917 ° W, 10 Jul 1932, T.A. Brindley, 13 (AMNH_PBI 00102479) (AMNH). 13 (AMNH_PBI 00107163), 1♀ (AMNH_PBI 00107189) (USNM); 05 Jun 1939 – 24 Jul 1939, T.A. Brindley, 33 (AMNH_ PBI 00107144–AMNH_PBI 00107146) (USNM); 21 Jul 1936, W.E. Schull, 1♀ (AMNH_PBI 00107179) (USNM). Twin Falls Co.: Hansen, 42.52963 ° N 114.3061 ° W, 01 Aug 1931, Harold Watersw, 1♀ (AMNH_PBI 00107172) (USNM). Unknown Co.: Ross, 14 Jul 1929, R.A. Flock, 13 (AMNH_PBI 00083532) (UCR). Montana: Fergus Co.: Lewistown, 47.06247 ° N 109.4282 ° W, 1204 m, 07 Jul 1961, G.M. Chamberlain, 1♀ (AMNH_PBI 00106288) (TAMU). Gallatin Co.: Beaver Creek, 44.85687 ° N 111.37356 ° W, 1920 m, 15 Aug 1913, S.J. Hunter, 1♀ (AMNH_PBI 00105192) (CNC). Bridger Range, 45.88861 ° N 110.9575 ° W, 1676 m, 10 Aug 1925, collector unknown, 13 (AMNH_PBI 00107152) (USNM). East Flathead Mont, 45.96935 ° N 110.89503 ° W, 1737 m, 25 Jul 1902, collector unknown, 1♀ (AMNH_PBI 00107183) (USNM). Moose Flat Campground, 26 mi S of Bozeman Hot Springs on Rt 191, 45.35527 ° N 111.17055 ° W, 1737 m, 10 Aug 1986, Schuh, Schwartz, and Stonedahl, 23 (AMNH_PBI 00102512, AMNH_PBI 00102513), 2♀ (AMNH_PBI 00102516, AMNH_PBI 00102517) (AMNH). Glacier Co.: Browning, 48.55694 ° N 113.0125 ° W, 26 Aug 1951, L.A. Konotopetz, 13 (AMNH_PBI 00105189) (CNC). Missoula Co.: Missoula, 46.87222 ° N 113.99306 ° W, 31 Jul 1920, A.A. Nichol, 13 (AMNH_PBI 00106284) (TAMU). 13 (AMNH_PBI 00107098), 2♀ (AMNH_PBI 00107106, AMNH_PBI 00107107) (USNM). Missoula County, 46.908 ° N 113.957 ° W, 29 Jun 1934, collector unknown, 13 (AMNH_PBI 00106285) (TAMU). Park Co.: 30 mi NW of Big Timber, 18 mi W of Route 191, Big Timber Campground, Crazy Mountains, Gallatin National Forest, 46.1413 ° N 110.39556 ° W, 2134 m, 01 Aug 1994, M.D. Schwartz, 13 (AMNH_ PBI 00105190) (CNC). Sanders Co.: Weeksville, 47.52299 ° N 114.994 ° W, 02 Aug 1882, collector unknown, 13 (AMNH_PBI 00102483), 2♀ (AMNH_PBI 00102487, AMNH_PBI 00102488) (AMNH). Nevada: Elko Co.: 16 mi SE of Elko on route 227, 40.72278 ° N 115.52125 ° W, 1707 m, 26 Jun 1983, R.T. Schuh and M.D. Schwartz, 1♀ (AMNH_PBI 00102528) (AMNH). Elko, 40.8325 ° N 115.76222 ° W, 12 Jul 1965, H.H. Knight, 13 (AMNH_PBI 00106616) (USNM). Esmeralda Co. : 12 mi W of Lida, 37.45813 ° N 117.71642 ° W, 08 Jul 1966, L. and C.W. O’Brien, 1♀ (AMNH_PBI 00104806) (CNC). 13 (AMNH_PBI 00105587) (TAMU). 13 (AMNH_PBI 00106617), 1♀ (AMNH_PBI 00106620) (USNM). 13 mi W of Lida on Rt 3, 37.45833 ° N 117.73397 ° W, 1938 m, 12 Jul 1980, R.T. Schuh and G.M. Stonedahl, Sarcobatus vermiculatus (Hook.) Torr. (Sarcobataceae) , det. J. Grimes 1980, 233 (AMNH_PBI 00102594–AMNH_PBI 00102615, AMNH_ PBI 00102792), 21♀ (AMNH_PBI 00102616– AMNH_PBI 00102636) (AMNH); 13 Jul 1980, R.T. Schuh and G.M. Stonedahl, Sarcobatus vermiculatus (Hook.) Torr. (Sarcobataceae) , det. J. Grimes 1980, 153 (AMNH_PBI 00102073–AMNH_PBI 00102087), 9♀ (AMNH_PBI 00102088–AMNH_PBI 00102095, AMNH_PBI 00102753) (AMNH); 12 Jul 1980, G.M. Stonedahl, Artemisia sp. (Asteraceae) , 23 (AMNH_PBI 00102569, AMNH_PBI 00102570), 4♀ (AMNH_PBI 00102574– AMNH_PBI 00102577) (AMNH); Sarcobatus vermiculatus (Hook.) Torr. (Sarcobataceae) , 183 (AMNH_PBI 00102161– AMNH_PBI 00102174, AMNH_PBI 00102775, AMNH_PBI 00102793), 20♀ (AMNH_PBI 00102176–AMNH_PBI 00102195), 13, 1♀ (in copula) (AMNH_PBI 00102175) (AMNH). Humboldt Co.: 18 mi N of Paradise Valley, Lye Creek Camp, 41.75443 ° N 117.53361 ° W, 2286 m, 07 Jul 1966, F., P., and B. Rindge, 2♀ (AMNH_PBI 00102518, AMNH_PBI 00102524) (AMNH). Lander Co.: 11 mi S of route 50 on route 376, T17N R44E, 39.25817 ° N 117.01958 ° W, 1768 m, 28 Jun 1983, R.T. Schuh and M.D. Schwartz, Sarcobatus vermiculatus (Sarcobataceae) , 13 (AMNH_PBI 00102250) (AMNH). Lyon Co.: 8 mi N of Sweetwater summit on highway 22, Toiyabe National Forest, 38.63046 ° N 119.26097 ° W, 1811 m, 11 Jul 1980, G.M. Stonedahl, Sarcobatus vermiculatus (Sarcobataceae) , 1♀ (AMNH_PBI 00102589) (AMNH). Nye Co.: 28 mi N of Belmont on Rt 82, 38.99273 ° N 116.70091 ° W, 2012 m, 13 Jul 1980, R.T. Schuh, Sarcobatusvermiculatus ( Sarcobataceae ), 13 (AMNH_PBI 00102249), 1♀ (AMNH_PBI 00102284) (AMNH); 13 Jul 1980, G.M. Stonedahl, Sarcobatus vermiculatus (Sarcobataceae) , 13 (AMNH_PBI 00102573), 1♀ (AMNH_PBI 00102588) (AMNH). White Pine Co.: 17.5 mi N of Hwy 50 on Steptoe Creek road, 39.39531 ° N 114.8324 ° W, 2341 m, 19 Jul 1980, G.M. Stonedahl, 1♀ (AMNH_ PBI 00102527) (AMNH). 31.8 mi N of Highway 50 on Steptoe Creek Road, 39.6806 ° N 114.75796 ° W, 1981 m, 19 Jul 1980, G.M. Stonedahl, Sarcobatus vermiculatus (Sarcobataceae) , 33 (AMNH_PBI 00102571, AMNH_ PBI 00102572, AMNH_PBI 00102774), 10♀ (AMNH_PBI 00102578–AMNH_PBI 00102587) (AMNH). New Mexico: San Juan Co.: 10 mi E of Blanco, 36.72403 ° N 107.64378 ° W, 14 Jun 1992, W.F. Chamberlain, 13 (AMNH_PBI 00105613) (TAMU). North Dakota: Billings Co.: 8 mi W of Belfield at Fryburg Oil Field, 46.84990N 103.40000W, 28 Jun 2000, T.J. Henry, Artemisia tridentata Nutt. (Asteraceae) , 23 (AMNH_PBI 00165796, AMNH_ PBI 00165797), 2♀ (AMNH_PBI 00165800, AMNH_PBI 00165801) (USNM). Slope Co.: 1 mi N of Marmarth, 46 ° 18 9 N 103 ° 93 9 W, 26 Jun 2000, T.J. Henry, Artemisia tridentata Nutt. (Asteraceae) , 23 (AMNH_PBI 00165798, AMNH_PBI 00165799), Symphoricarpos occidentalis Hook. (Caprifoliceae) , 43 (AMNH_ PBI 00165792–AMNH_PBI 00165795), 1♀ (AMNH_PBI 00165802) (USNM). Oregon: Benton Co.: Corvallis, 44.56472 ° N 123.26083 ° W, 26 Jun 1926, C.J. Drake, 1♀ (AMNH_PBI 00107177) (USNM); 09 Jul 1947, H.H. Crowell, seed cabage, 23 (AMNH_PBI 00107165, AMNH_PBI 00107166) (USNM). Columbia Co. : Scappoose, 45.75444 ° N 122.87639 ° W, 16 Jul 1935, K. Gray, 13 (AMNH_PBI 00102481) (AMNH). Crook Co.: 3 mi S of Prineville, 44.25728 ° N 120.85429 ° W, 02 Aug 1935, J. Schuh, 23 (AMNH_PBI 00102247, AMNH_PBI 00102248) (AMNH). Harney Co.: Albert Lake, 43.44959 ° N 119.1377 ° W, 02 Jul 1935, J. Schuh, 13 (AMNH_PBI 00102482) (AMNH). Hood River Co.: Hood River, 45.71556 ° N 121.51 ° W, Jul 1931, R.H. Beamer, 33 (AMNH_PBI 00075258, AMNH_ PBI 00075259, AMNH_PBI 00075279), 1♀ (AMNH_PBI 00075276) (KU). Mount Hood, 45.53806 ° N 121.56722 ° W, 03 Jul 1935, R.H. Beamer, 1♀ (AMNH_PBI 00075265) (KU). Jackson Co.: 0.5 mi S of Siskiyou Summit on Old Rt 99, Old Siskiyou Rd, 42.06777 ° N 122.60583 ° W, 1311 m, 27 Jun 1979, M.D. Schwartz, Lupinus sp. (Fabaceae) , 13 (AMNH_PBI 00102466) (AMNH). 0.5 mi S of Siskiyou Summit on Old Rt 99, Old Siskiyou Rd, 42.06777 ° N 122.60583 ° W, 1346 m, 26 Jun 1979, R.T. and Joe Schuh, Lupinus sp. (Fabaceae) , 13 (AMNH_PBI 00102457), 1♀ (AMNH_PBI 00102475) (AMNH). Shady Cove, 42.61068 ° N 122.8125 ° W, 427 m, 23 Jun 1977, J. Schuh, 13 (AMNH_PBI 00102463), 3♀ (AMNH_PBI 00102472–AMNH_ PBI 00102474) (AMNH). Siskiyou Summit, Old Road, 42.075 ° N 122.60583 ° W, 1260 m, 22 Jul 1999, Schwartz, Gillespie, Quiring, Lupinus sp. (Fabaceae) , 4♀ (AMNH_PBI 00103899– AMNH_PBI 00103902) (CNC); 28 Jul 1999, M.D. Schwartz, Chrysothamnus nauseosus (Pall. ex Pursh) Britt. (Asteraceae) , 13 (AMNH_PBI 00103896) (CNC). Siskiyou Summit, Old Siskiyou Hwy and Frontage Rd, 42.075 ° N 122.60583 ° W, 1314 m, 01 Jul 1994, M.D. Schwartz, Lupinus sp. (Fabaceae) , 13 (AMNH_PBI 00105188), 1♀ (AMNH_ PBI 00105191) (CNC). ust E of Pinehurst, 42.11778 ° N 122.365 ° W, 1340 m, 27 Jun 1979, R.T. and Joe Schuh, 23 (AMNH_PBI 00102459, AMNH_PBI 00102460), 2♀ (AMNH_ PBI 00102477, AMNH_PBI 00102478) (AMNH). Josephine Co.: 12 mi N of Cave Junction, 42.33654 ° N 123.64694 ° W, 470 m, 10 Jul 1979, R.T. and Joe Schuh, 13 (AMNH_ PBI 00102458), 1♀ (AMNH_PBI 00102476) (AMNH). Lake Co.: 11 mi N of Silver Lake, 43.25913 ° N 120.888 ° W, 1500 m, 25 Jun 1979, R.T. Schuh, 23 (AMNH_PBI 00102470, AMNH_PBI 00102471) (AMNH). Lane Co.: Blue River Reservoir, 44.15472 ° N 122.34 ° W, 31 Jul 1979, G.M. Stonedahl, Chrysolepis chrysophylla (Fagaceae) , 13 (AMNH_PBI 00102467) (AMNH). Linn Co.: HJ Andrews Experimental Forest, 1 mile from Road 350 end, Ridge Site, 44.2325 ° N 122.1167 ° W, 16 Jul 1985, Stonedahl & McIver, Lupinus sp. (Fabaceae) , 13 (AMNH_ PBI 00102495), 1♀ (AMNH_PBI 00102496) (AMNH). HJ Andrews Experimental forest, Carpenter Mountain, end of Natl Forest Rd 1506-350, 44.27834 ° N 122.14628 ° W, 1448 m, 30 Aug 1978, G.M. Stonedahl, Anaphalis margaritacea (Asteraceae) , 13 (AMNH_PBI 00102497), 2♀ (AMNH_PBI 00102498, AMNH_ PBI 00102499) (AMNH). Umatilla Co.: Meacham, 45.50667 ° N 118.42028 ° W, 14 Jul 1931, H.T. Peters, 23 (AMNH_PBI 00075255, AMNH_PBI 00075256) (KU); 14 Jul 1931, R.H. Beamer, 13 (AMNH_PBI 00075254) (KU). Union Co.: 6 mi NW of Medical Springs, summit of Wallowa Mountains, Wallowa-Whitman National Forest T5S R41E Sec27, 45.017 ° N 117.6758 ° W, 1073 m, 29 Jul 1994, M.D. Schwartz, Lupinus leucophyllus Dougl. (Fabaceae) , 13 (AMNH_PBI 00103895), 2♀ (AMNH_PBI 00103897, AMNH_ PBI 00103898) (CNC). Elgin, 45.56486 ° N 117.9174 ° W, 21 Jul 1929, H.A. Scullen, Monardella villosa (Lamiaceae) , 13 (AMNH_PBI 00107169) (USNM). La Grande, 45.32472 ° N 118.08667 ° W, 26 Jun 1926, E.W. Davis, 23 (AMNH_PBI 00107094, AMNH_PBI 00107095), 3♀ (AMNH_PBI 00107101–AMNH_PBI 00107103) (USNM). Unknown Co.: Dixie, 08 Jul 1931, J. Nottingham, 1♀ (AMNH_PBI 00075263) (KU). Long creek, 01 Jul 1935, J. Schuh, 13 (AMNH_PBI 00102480) (AMNH). South Dakota: Lawrence Co.: Black Hills, 44.41667 ° N 103.70833 ° W, 28 Jun 1937, H.M. Harris, 1♀ (AMNH_PBI 00107204) (USNM). Deadwood, 44.37667 ° N 103.72917 ° W, 29 Jul 1927, H.H. Knight, 13 (AMNH_PBI 00107092), 2♀ (AMNH_PBI 00107099, AMNH_PBI 00107100) (USNM). Meade Co.: Piedmont, 44.23165 ° N 103.389 ° W, 1065 m, 17 Jul 1937, C.L. Johnston, 1♀ (AMNH_PBI 00075278) (KU). Utah: Box Elder Co. : 5 mi SW of Clear Creek Campground, Raft River Mountains, 41.94517 ° N 113.35486 ° W, 2164 m, 31 Jul 1981, M.D. Schwartz, Lupinus sp. (Fabaceae) , 1♀ (AMNH_PBI 00102525) Holodiscus discolor (Rosaceae) , 13 (AMNH_PBI 00102469) (AMNH). Raft River Mountains, 5 mi SW Clear Crk. Cmpgrd. T14N R13N, 41.99118 ° N 113.64017 ° W, 2164 m, 31 Jul 1981, M.D. Schwartz, Lupinus sp. (Fabaceae) , 13 (AMNH_PBI 00102777) (AMNH). Cache Co.: Wellsville, 41.63861 ° N 111.93306 ° W, 17 Jun 1940, G.F. Knowiten and M.J. Janes, 13 (AMNH_PBI 00107168) (USNM). SanJuan Co.: Geyser Creek, near Taylor Flats, 38.49469 ° N 109.09081 ° W, 04 Jul 1980, J.T. and D.A. Polhemus, 2♀ (AMNH_PBI 00065090, AMNH_ PBI 00065091) (JTP). Sanpete Co.: 13 mi E of Fairview on Rt 31, T14S R6E, 39.61952 ° N 111.28186 ° W, 2286 m, 08 Aug 1981, M.D. Schwartz, Conyza canadensis (L.) Cronq. ( Asteraceae ), 13 (AMNH_PBI 00102468) (AMNH). Sevier Co.: Monroe, 38.63 ° N 112.12 ° W, 25 Jul 1906, collector unknown, 1♀ (AMNH_PBI 00106614) (USNM). Summit Co.: 4.5 mi E of Oakley, Weber Canyon Road, 40.71472 ° N 111.21417 ° W, 19 Aug 1986, G.M. Stonedahl, Cercocarpus sp. (Rosaceae) , 1♀ (AMNH_PBI 00102529) (AMNH). Uintah Co.: 5–10 mi SW of Bonanza, T10S R24E Sec 17 (R2), 39.9456 ° N 109.2406 ° W, 1615 m, 08 Jul 1982, M.D. Schwartz, Sarcobatus vermiculatus (Sarcobataceae) , 1♀ (AMNH_PBI 00102418) (AMNH). 5–10 mi SW of Bonanza, T10S R24E Sec 17 (R4), 39.9456 ° N 109.2406 ° W, 1615 m, 05 Jul 1982, M.D. Schwartz, Sarcobatus vermiculatus (Sarcobataceae) (Hook.) Torr. ( Sarcobataceae ), 83 (AMNH_PBI 00102054, AMNH_PBI 00102408–AMNH_PBI 00102412, AMNH_ PBI 00102776, AMNH_PBI 00102794), 9♀ (AMNH_PBI 00102413–AMNH_PBI 00102417, AMNH_PBI 00102419–AMNH_PBI 00102421, AMNH_PBI 00102756) (AMNH). Weber Co.: Weber Canyon, 41.13578 ° N 111.90382 ° W, 04 Jul 1931, M.W. Sanderson, 1♀ (AMNH_PBI 00075264) (KU). Unknown Co.: Dry Lake, 25 Jul 1938, G.F. Knowlton, 1♀ (AMNH_PBI 00107174) (USNM). Washington: Asotin Co.: 2.5 mi S of Anatone, 2 mi N Rattlesnake Summit, 46.09885 ° N 117.13139 ° W, 1189 m, 04 Aug 1986, Schuh, Schwartz, and Stonedahl, Lupinus sp. (Fabaceae) , 13 (AMNH_ PBI 00102644) (AMNH). Chelan Co.: Leavenworth, 47.59623 ° N 120.6614 ° W, 07 Jul 1933 – 09 Jul 1933, Wickham, 13 (AMNH_ PBI 00107129) (USNM). Wenatchee, 47.42345 ° N 120.3103 ° W, 14 Jun 1915, E.J. Newcomer, 1♀ (AMNH_PBI 00107173) (USNM). Columbia Co. : Dayton, 46.32389 ° N 117.97139 ° W, 1447 m, 06 Jul 1937, R.E. Miller, 23 (AMNH_PBI 00107096, AMNH_PBI 00107097), 2♀ (AMNH_PBI 00107104, AMNH_PBI 00107105) (USNM). Cowlitz Co.: Kalama R[iver], 46.03372 ° N 122.87038 ° W, 21 Jul 1931, R.H. Beamer, 13 (AMNH_PBI 00075260) (KU). Ferry Co.: Republic, 48.64833 ° N 118.73667 ° W, 06 Aug 1931, H.T. Peters, 13 (AMNH_PBI 00075257), 1♀ (AMNH_PBI 00075277) (KU). Kittitas Co.: Cliffdell, 46.94639 ° N 121.06778 ° W, 07 Jul 1935, R.H. Beamer, 13 (AMNH_PBI 00075253) (KU). Okanogan Co.: 8 mi WNW of Republic (Ferry Co.), Sweat Creek, 48.42324 ° N 119.83129 ° W, 1097 m, 20 Jul 1978, N. Herman, 13 (AMNH_PBI 00102465) (AMNH). Pierce Co.: Mount Rainier, cotton fields, 46.8833 ° N 122.88333 ° W, 07 Jul 1935, Oman, 1♀ (AMNH_PBI 00107134) (USNM). Whitman Co.: Pullman, 46.73139 ° N 117.17861 ° W, 29 Aug 1909, J.A. Hyslop, 1♀ (AMNH_PBI 00107175) (USNM); 14 Jul 1911, collector unknown, 13 (AMNH_PBI 00107124) (USNM); 24 Jul 1908, W.M. Mann, 13 (AMNH_PBI 00107125), 2♀ (AMNH_PBI 00107131, AMNH_PBI 00107133) (USNM); 30 Jul 1909, J.A. Hyslop, 23 (AMNH_ PBI 00107126, AMNH_PBI 00107128) (USNM); 17 Jul 1908, W.M. Mann, 13 (AMNH_PBI 00107127) (USNM); 08 Jul 1908, W.M. Mann, 1♀ (AMNH_ PBI 00107132) (USNM); 02 Aug 1908, W.M. Mann, 13 (AMNH_PBI 00107153) (USNM); 02 Aug 1908, W.M. Mann, 13 (AMNH_PBI 00105186) (CNC). Yakima Co.: 10 mi N of Yakima, 46.73297 ° N 120.45236 ° W, 04 Jul 1978, J. Schuh & R. Rieder, 13 (AMNH_PBI 00102464) (AMNH). Naches, 46.73111 ° N 120.69833 ° W, 07 Jul 1935, R.H. Beamer, 1♀ (AMNH_PBI 00075262) (KU). Tieton Canyon, 46.65217 ° N 120.7248 ° W, 21 Jun 1932, A.R. Rolfs, 13 (AMNH_PBI 00106286) (TAMU). Toppenish, 46.3775 ° N 120.3075 ° W, 12 Jun 1926, E.W. Davis, 13 (AMNH_PBI 00107157) (USNM); 15 May 1926, E.W. Davis, 1♀ (AMNH_PBI 00107185) (USNM). Yakima, 46.60222 ° N 120.50472 ° W, 11 Aug 1932, A.R. Rolfs, 13 (AMNH_PBI 00106287) (TAMU); 21 Aug 1931, A.R. Rolfs, 13 (AMNH_PBI 00107160), 1♀ (AMNH_PBI 00107187) (USNM); 01 May 1931, A.R. Rolfs, 13 (AMNH_ PBI 00107161) (USNM). Wyoming: Albany Co.: 40 mi NE of Laramie, 41.80783 ° N 105.28074 ° W, 13 Jul 1937, R.H. Beamer, 53 (AMNH_PBI 00075267–AMNH_PBI 00075271), 4♀ (AMNH_PBI 00075280–AMNH_PBI 00075283) (KU). Big Horn Co.: Big Horn Mountains, 44.75499 ° N 107.78239 ° W, 1981 m, 17 Aug 1927, H.H. Knight, 1♀ (AMNH_PBI 00107182) (USNM). Crook Co.: Sundance, 44.40639 ° N 104.37528 ° W, 30 Jul 1927, H.H. Knight, 13 (AMNH_PBI 00105187) (CNC). 73 (AMNH_PBI 00107085–AMNH_PBI 00107091), 7♀ (AMNH_PBI 00107112–AMNH_PBI 00107116, AMNH_PBI 00107118, AMNH_PBI 00107119) (USNM). Fremont Co.: Shoshone National Forest, 43.80913 ° N 109.56306 ° W, 14 Aug 1927, H.H. Knight, 13 (AMNH_PBI 00125480) (CUIC); 15 Aug 1927, H.H. Knight, 13 (AMNH_PBI 00107302) (USNM). Johnson Co. : Fort McKinley, 44.33932 ° N 106.73531 ° W, Jul 1883, collector unknown, 13 (AMNH_PBI 00107167) (USNM). Lincoln Co.: Salt River Pass, 15 mi S of Afton on Rt 89, 42.54234 ° N 110.89431 ° W, 2326 m, 21 Jul 1981, M.D. Schwartz, Artemisia tridentata (Asteraceae) , 1♀ (AMNH_PBI 00102526) Lupinus sp. (Fabaceae) , 13 (AMNH_PBI 00102764) (AMNH). Snake River, 18 mi E Alpine, 43.23 ° N 110.77 ° W, 12 Aug 1949, W.J. and J.W. Gertsch, 13 (AMNH_PBI 00102795) (AMNH). Park Co.: 19 mi E of Cooke City on Rt 212, 44.93593 ° N 109.62787 ° W, 2438 m, 11 Aug 1986, Schuh, Schwartz, and Stonedahl, Lupinus sp. (Fabaceae) , 13 (AMNH_ PBI 00102511), 1♀ (AMNH_PBI 00102515) (AMNH). National Park [Yellowstone], 44.76667 ° N 110.23333 ° W, collector unknown, 1♀ (AMNH_PBI 00107176) (USNM); O. Heidemann, 13 (AMNH_PBI 00125481), 1♀ (AMNH_PBI 00125483) (CUIC); 26 Jul 1891, collector unknown, 13 (AMNH_PBI 00125482) (CUIC); 26 Jul 1891, O. Heidemann, 2♀ (AMNH_PBI 00125484, AMNH_PBI 00125485) (CUIC). Shoshone National Forest, 44.00028 ° N 109.50028 ° W, 07 Aug 1927, H.H. Knight, 23 (AMNH_PBI 00107083, AMNH_PBI 00107084) (USNM); 14 Aug 1927, H.H. Knight, 1♀ (AMNH_ PBI 00106290) (TAMU). 193 (AMNH_PBI 00107064–AMNH_PBI 00107082), 5♀ (AMNH_ PBI 00107108–AMNH_PBI 00107111, AMNH_PBI 00107117) (USNM). Sunlight Creek, 44.69467 ° N 109.82018 ° W, 31 Jul 1896, R.P. Currie, 1♀ (AMNH_PBI 00107149) (USNM). Yellowstone National Park, 44.76667 ° N 110.23333 ° W, 08 Aug 1927, H.H. Knight, 13 (AMNH_PBI 00107162), 1♀ (AMNH_PBI 00107188) (USNM).Yellowstone National Park, 44.76667 ° N 110.23333 ° W, 04 Aug 1931, H.H. Knight, 13 (AMNH_PBI 00107301), 1♀ (AMNH_PBI 00107178) (USNM). Sheridan Co.: SW of Sheridan, 44.79722 ° N 106.95556 ° W, 16 Aug 1979, J.T. Polhemus, 1♀ (AMNH_ PBI 00065103) (JTP). Sweetwater Co.: Green River Lake, 41.52299 ° N 109.46945 ° W, 2591 m, 01 Aug 1935, H. Ruckes, 13 (AMNH_ PBI 00107196), 1♀ (AMNH_PBI 00107202) (USNM). Teton Co.: Grand Teton National Park, 2 mi SE Colter Bay Village, St. 89, 43.87622 ° N 110.61996 ° W, 2134 m, 22 Jul 1981, M.D. Schwartz, Lupinus sp. (Fabaceae) , 43 (AMNH_PBI 00101208, AMNH_ PBI 00102500–AMNH_PBI 00102502), 8♀ (AMNH_PBI 00102503–AMNH_PBI 00102510) (AMNH). Stewart R. Sta., F 4749, 43.7 ° N 110.75 ° W, 2042 m, 18 Jul 1920, collector unknown, 13 (AMNH_PBI 00102461) (AMNH). Unknown state: Unknown Co.: Olmstead, 13 (AMNH_PBI 00102484) (AMNH). Western Territories, 13 (AMNH_PBI 00107130), 2♀ (AMNH_PBI 00107140, AMNH_PBI 00107141) (USNM).