Pupina arula Benson, 1856

Jirapatrasilp, Parin, Sutcharit, Chirasak & Panha, Somsak, 2022, Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia, ZooKeys 1119, pp. 1-115 : 1

Pupina arula Benson, 1856

Figs 29A, B , 33D

Pupina arula Benson, 1856: 230. Type locality: ad Yunglaw, in valle Tenasserim [Tanintharyi Region, Myanmar]. Theobald 1858 [1857]: 247. Pfeiffer 1860: 141, pl. 37, figs 7-9. Hanley and Theobald 1870: 4, pl. 7, fig. 4. Reeve 1878: Pupinidae , pl. 1, sp. 5. Crosse 1879: 340 (part). de Morgan 1885: 413 (part). von Möllendorff 1887 [1886]: 314 (part). Godwin-Austen 1897: 37, 38, pl. 69, fig. 1, 1a. BEDO 2017: 88. Sutcharit et al. 2018: fig. 5-13e.

Pupina avula [sic]-Sowerby I 1866: Pupinidae , pl. 3 (pl. 265), Pupina , fig. 3.

Pupina (Tylotoechus) arula - Kobelt 1902: 307. Gude 1921: 193, 194 (part). Solem 1966: 12, Doi Sutep [Doi Suthep Mountain, Chiang Mai Province, Thailand]. Hemmen and Hemmen 2001: 39.

Pupina arula arula - Maassen 2001: 40.

Type material examined.

Syntype UMZC I.103025 (1 shell; Figs 29A , 33D ) from the R. McAndrew collection labelled "Bens. col., Ind".

Other material examined.

Specimen NHMUK 1888.12.4.109 (1 shell; Fig. 29B ) from Yunglaw, Myanmar, the W. Theobald collection.

Diagnosis.

Shell ovate; last whorl ca. 80% of shell height. Shell surface matt. Apertural lip thickened but not expanded; apertural lip curved when observed from lateral view. Columellar tooth fin-shaped.

Differential diagnosis.

Pupina arula can be distinguished from all other species in the P. arula species group from mainland Southeast Asia by a matt shell surface.

Distribution.

Myanmar and an uncertain record from northern Thailand ( Solem 1966).

Remarks.

No material of this species was found during this survey. The specimen of P. arula mentioned in Davison (1995: 237) from Temengor dam, Perak, Malaysia possibly belongs to P. perakensis .

Figures

Figure 29. Shells of Pupina arula species group from mainland Southeast Asia A, B Pupina arula A syntype UMZC I. 103025 and B specimen NHMUK 1888.12.4.109. C, D Pupina mouhoti, possible syntypes NHMUK ex. Cuming coll. E, F Pupina vescoi E syntype NHMUK 1893.2.4.767 and F specimen SMF 109956 / 1 G Pupina perakensis, lectotype SMF 109969 / 1 H Pupina excisa, lectotype SMF 110778 / 1 I-K Pupina exclamationis I, J syntypes MNHN-IM- 2000 - 35840 and K specimen NHMUK 1901.12.23.205 " forma minor ". Photo: J. Ablett, H. Taylor, NHM (A), A. Lardeur, P. Maestrati, MNHN (I, J).

Figure 33. A Pupina bilabiata sp. nov., specimen CUMZ 12086 from Na San Temple, Surat Thani B, C Pupina godwinausteni sp. nov. B holotype CUMZ 12090 / 1 and C paratype CUMZ 12090 / 2 from Khao Wong Cave, Uthai Thani D Pupina arula, syntype UMZC I. 103025 " Ind " E Pupina mouhoti, possible syntype NHMUK ex. Cuming coll. from Camboja F Pupina vescoi, syntype NHMUK 1893.2.4.767 from Cochin China. Photo: J. Ablett, H. Taylor, NHM (D).