Rhinusa (Caldara, 2001)

Caldara, Roberto & Toševski, Ivo, 2019, Rhinusa Stephens: a taxonomic revision of the species belonging to the R. linariae R. herbarum, R. melas, and R. mauritii groups (Coleoptera Curculionidae), Zootaxa 4679 (2), pp. 318-340 : 338

Key to all the groups and treated species of Rhinusa

1. Protibiae and mesotibiae with premucro, more pronounced in female ( Figs 25 and 26 ); in male protibiae distinctly arcuate in apical quarter, and profemora distinctly swollen with distinct tooth ( Fig. 25 ); ventrites 3–5 in male along midline with dense and ruffled hairlike scales..................................................................... R. tetra group

- Protibiae and mesotibiae without premucro in both sexes ( Figs 23 and 24 ); in male protibiae at most moderately arcuate in apical quarter, and profemora less swollen at most with small tooth ( Figs 23 and 24 ); ventrites 3–5 in male along midline without hairlike scales........................................................................................ 2

2. Rostrum in dorsal view from base to antennal insertion with a trapezoidal outline, with dorsal part narrower than ventral part. Rostrum of male in dorsal view with clearly visible scrobes ( Fig. 22 ).............................. R. antirrhini group

- Rostrum otherwise.................................................................................... 3

3. Scales of dorsal vestiture short, oval to elliptical ( Fig. 9 ).............................................. R. mauritii

- Scales of dorsal vestiture long, setiform ( Figs 1–8 )........................................................... 4

4 Meso- and metafemora at most with a small tooth similar to that on profemora..................................... 5

- Meso- and metafemora with distinct sharp tooth, usually larger than that on profemora............................. 14

5. Dorsal vestiture composed of erect very long setiform scales, at least 20× as long as wide................. R. pilosa group

- Dorsal vestiture composed of recumbent and suberect setiform scales, at most 12× as long as wide..................... 6

6. Tibiae with outer margin distinctly curved outward at apex ( Fig. 24 ). Rostrum in lateral view strongly arcuate ( Figs 10 and 11 )................................................................................... R. linariae group 7

- Tibiae with outer margin curved inward near apex ( Fig. 23 ). Rostrum in lateral view weakly to moderately curved........ 9

7. Elytra and legs black ( Fig. 1 )..................................................................... R. linariae

- Elytra and legs reddish ( Figs 2 and 3 )..................................................................... 8

8. Integument completely reddish. Elytra longer, subelliptical (El/Ew 1.20–1.24). Pronotum less transverse (Pw/Pl 1.37–1.40) ( Fig. 3 ). Outer margin of tibiae weakly carinate.................................................. R. kumatschevi

- Integument partly blackish. Elytra short, suboval (El/Ew 1.10–1.14). Pronotum more transverse (Pw/Pl 1.44–1.49) ( Fig. 2 ). Outer margin of tibiae strongly carinate............................................................. R. brisouti

9. Elytra short (El/Ew <1.20) ( Figs 7 and 8 ). Rostrum poorly sexually dimorphic and short in both sexes ( Figs. 16–19 )............................................................................................. R. melas group 10

- Elytra elongate (El/Ew> 1.30) ( Figs 4–6 ). Rostrum in female distinctly different in shape and moderately to distinctly longer than in male (except for R. mateui , but in this case rostrum distinctly thinner in both sexes, figs 14–15)................ 11

10. Legs blackish. Scales of dorsal vestiture greyish without sericeous reflection ( Fig. 7 ). Tarsomere 3 distinctly bilobed and distinctly wider than tarsomere 2..................................................................... R. melas

- Legs reddish. Scales of dorsal vestiture greyish with distinct sericeous reflection ( Fig. 8 ). Tarsomere 3 weakly bilobed and moderately wider than tarsomere 2.............................................................. R. korotyaevi

11. Rostrum in lateral view distinctly tapered from base to apex ( Figs 12 and 13 )..................................... 12

- Rostrum in lateral view of subequal width from base to apex ( Figs 15 and 16 ).................................... 13

12. Pronotum more transverse (Pw/Pl 1.34–1.37) and with rounded sides. Elytral integument reddish ( Fig. 4 )...... R. herbarum

- Pronotum less transverse (Pw/Pl 1.28), subconical, with weakly rounded sides. Elytral integument blackish ( Fig. 5 )................................................................................................. R. vulpeculus

13. Tarsomere 3 weakly bilobed and slightly wider than tarsomere 2. Elytral integument blackish. Length 1.3–1.6 mm ..................................................................................................... R. mateui

- Tarsomere 3 distinctly bilobed and distinctly wider than tarsomere 2. Elytral integument usually partly reddish. Length 1.5–3.5 mm ................................................................................. R. bipustulata group

14. Rostrum stout in both sexes, in female in lateral view gradually tapered from antennal insertion to apex, usually shorter (Rl/Pl in male <0.75, in female <0.85). Elytra with distinctly pronounced humeri, distinctly flattened on disc...... R. vestita group

- Rostrum slender in both sexes, especially in female where in lateral view it is cylindrical and of equal width from antennal insertion to apex, usually longer (Rl/Pl in male> 0.80, in female> 0.90). Elytra with weakly pronounced humeri, slightly to moderately convex on disc.................................................................... R. neta group

Figures

FIGURES 23‒32. Profemur and protibia of (23) Rhinusa bipustulata, male; (24) R. linariae, male; (25) R. tetra, male; (26) R. tetra, female. Penis in dorsal view of (27) R. linariae; (28) R. herbarum; (29) R. mateui; (30) R. melas; (31) R. korotyaevi; (32) R. mauritii.

FIGURES 10‒22. Rostrum of (10) Rhinusa linariae, male; (11) R. linariae, female; (12) R. herbarum, male; (13) R. herbarum, female; (14) R. mateui, male; (15) R. mateui, female; (16) R. melas, male; (17) R. melas, female; (18) R. korotyaevi, male; (19) R. korotyaevi, female; (20) R. mauritii, male; (21) R. mauritii, female; (22) R. antirrhini, male.

FIGURES 1‒9. Habitus of (1) Rhinusa linariae, male; (2) R. brisouti, male; (3) R. kumatschevi, male; (4) R. herbarum, male; (5) R. vulpeculus, holotype male; (6) R. mateui, female; (7) R. melas, male; (8) R. korotyaevi, paratype male; (9) R. mauritii, male. Not to the same scale