Trichopalpus Rondani, 1856

Ozerov, A. L. & Krivosheina, M. G., 2023, Contribution to the fauna of dung flies (Diptera: Scathophagidae) of Russia with a key to genera and a checklist of the Russian Scathophagidae, Russian Entomological Journal 32 (1), pp. 95-122 : 119-121

Genus Trichopalpus Rondani, 1856

Figs 20 , 23, 27 , 127, 128 .

REMARKS. Diagnosis of the genus, descriptions of the species noted in Russia, a key for their identification and changes in nomenclature are given by Ozerov [2019c]. Currently, there are 6 species in Russia (see checklist below).

Checklist of the family Scathophagidae of Russia

Acanthocnema Becker, 1894

altaica Ozerov et Krivosheina, 2018

latipennis Becker, 1894

proboscidea Ozerov et Krivosheina, 2018

sternalis Suwa, 1986

=vikhrevi Ozerov et Krivosheina, 2014

Acerocnema Becker, 1894

arctica Ozerov, 2013

barkalovi Ozerov, 2007

breviseta (Zetterstedt, 1846 — Cordylura )

=tiefi Becker, 1894

gorodkovi Ozerov et Krivosheina, 2018

lobanovi Ozerov, 2007

macrocera (Meigen, 1826 — Cordylura )

nitens Ozerov et Krivosheina, 2018

paradoxopyga Stackelberg, 1952

richterae Ozerov, 2015

Bostrichopyga Becker, 1894

crassipes (Zetterstedt, 1838 — Cordylura )

Cleigastra Macquart, 1835

= Gonatherus Rondani, 1856

= Gonarcticus Becker, 1894

= Nanna Strobl, 1894

= Amaurosoma Becker, 1894

= Spathephilus Becker, 1894

= Orthacheta Becker, 1894

abdominalis (Zetterstedt, 1846 — Cordylura )

amurensis (Ozerov, 2010 — Nanna )

antennata (Zetterstedt, 1838 — Scatomyza )

apicalis (Meigen, 1826 — Cordylura )

arctica (Becker, 1907 — Pselaphephila )

armillata (Zetterstedt, 1846 — Cordylura )

articulata ( Becker, 1894 — Amaurosoma )

bispinosa (Malloch, 1920 — Amaurosoma )

brevifrons (Zetterstedt, 1838 — Cordylura )

cornuta (Loew, 1863 — Cordylura )

cryophila ( Ozerov et Krivosheina, 2015 — Nanna )

flavipes ( Fallén, 1819 — Cordylura )

inermis ( Becker, 1894 — Amaurosoma )

intermedia Ozerov et Krivosheina, 2016

katmaiensis (Malloch, 1920 — Amaurosoma )

leucostoma (Zetterstedt, 1846 — Cordylura )

loewi ( Becker, 1894 — Pselaphephila )

longicornis (von Roser, 1840 — Cordylura )

maritima Ozerov et Krivosheina, 2016

nigriventris (Loew, 1864 — Cordylura )

obscuripes (Becker, 1915 — Acerocnema )

=indotatum (Engelmark, 1999 — Nanna )

pilosa (Zetterstedt, 1838 — Cordylura )

planiceps (Fallén, 1826 — Cordylura )

=fumipennis (Hendel, 1930 — Gonatherus )

puberula ( Becker, 1894 — Amaurosoma )

rossolimoae (Ozerov, 2010 — Nanna )

sundukovi Ozerov, 2013

tibiella (Zetterstedt, 1838 — Cordylura )

Cordilura Fallén, 1810

= Phrosia Robineau-Desvoidy, 1830

= Scoliaphleps Becker, 1894

= Cordilurina James, 1955

= Cordylura , unjustified emend.

aberrans ( Becker, 1894 — Cordylura )

= rubifrontata Becker, 1894 — Cordylura ).

aemula (Collín, 1958 — Cordylura )

albilabris (Fabricius, 1805 — Ocyptera )

albipes ( Fallén, 1819 — Cordylura )

amurensis Ozerov, 2007

atrata (Zetterstedt, 1846 — Cordylura )

bicoloripes Ozerov, 1997

ciliata (Meigen, 1826 — Cordylura )

fulvifrons Ozerov, 1997

fuscipes (Zetterstedt, 1838)

= apicata (Hendel, 1930 — Cordylura )

= nigrithorax (Hendel, 1930 — Cordylura )

= ochracea (Hendel, 1930 — Cordylura )

grunini Ozerov et Krivosheina, 2017

impudica (Rondani, 1867 — Cordylura )

= umbrosa (Loew, 1873 — Cordylura )

kakaberrans Ozerov, 1997

kosterini Ozerov et Krivosheina, 2014

krocha Ozerov, 2007

monochroma Ozerov et Krivosheina, 2014

nartshukae Ozerov et Krivosheina, 2015

negrobovi Ozerov et Krivosheina, 2017

nubecula Sasakawa, 1986

nubilosa Ozerov et Krivosheina, 2020

picipes (Meigen, 1826 — Cordylura )

picticornis (Loew, 1864 — Cordylura )

= latigenis Hendel, 1930 — Cordylura )

proboscidea (Zetterstedt, 1838 — Cordylura )

pudica (Meigen, 1826 — Cordylura )

remmi Elberg, 1972

remota Ozerov, 1997

richterae Ozerov et Krivosheina, 2015

rufimana (Meigen, 1826 — Cordylura )

rufipes (Meigen, 1826 — Cordylura )

= pubera Linnaeus, 1758 sensu Fallén, 1810:15 , misidentification sagittifera Gorodkov, 1974

shatalkini Ozerov, 1997

sibirica Gorodkov, 1974

sidorenkoi Ozerov et Krivosheina, 2012

sifneri Ozerov, 2007

socialis ( Becker, 1894)

tartariana Ozerov, 2007

ustulata (Zetterstedt, 1838 — Cordylura )

zaitzevi Gorodkov, 1974

Cosmetopus Becker, 1894

dentimanus (Zetterstedt, 1838 — Cordylura )

longus (Walker, 1849 — Cordylura )

=fulvipes (Zetterstedt, 1838 — Cordylura )

Delina Robineau-Desvoidy, 1830

nigrita ( Fallén, 1819 — Cordylura )

Ernoneura Becker, 1894

argus (Zetterstedt, 1838 — Scatomyza )

Gimnomera Rondani, 1867

= Cochliarium Becker, 1894

= Langechristia Ozerov, 1999

= Gymnomera , error alanica (Ozerov, 1999 — Cochliarium ) albipila (Zetterstedt, 1846 — Cordilura ) = lucida (Becker, 1900 — Microprosopa ) amica (Ozerov, 1999 — Langechristia ) castanipes ( Becker, 1894 — Cochliarium ) cuneiventris (Zetterstedt, 1846 — Cordilura ) dorsata (Zetterstedt, 1838 — Cordilura )

hirta Hendel, 1930 — Gymnomera ). lasiostoma ( Becker, 1894 — Cochliarium ) mellina (Becker, 1900 — Gymnomera ) montana Ozerov et Krivosheina, 2013 nigricorpus Ozerov, 2019

novgorodovae (Ozerov, 2007 — Cochliarium ) = sorokinae (Ozerov, 2007 — Cochliarium ) nudipedis Ozerov, 2019

palliseta Ozerov, 2019

speciosa (Ozerov, 1999 — Langechristia ) tarsea ( Fallén, 1819 — Cordilura )

tibialis (Malloch, 1919 — Dasypleuron ) = sibirica (Engelmark, 1999 — Cochliarium ) tukuringra (Ozerov, 1999 — Cochliarium ) ziegleri (Ozerov, 1999 — Langechristia )

Hexamitocera Becker, 1894

loxocerata (Fallén, 1826 — Cordilura )

Hydromyza Fallén, 1823

livens (Fabricius, 1794 — Musca )

Leptopa Zetterstedt, 1838

= Parallelomma Becker, 1894

= Chylizosoma Hendel, 1924

caucasica (Ozerov, 2010 — Parallelomma ) filiformis Zetterstedt, 1838

hostae (Herring, 1955 — Chylizosoma )

media ( Becker, 1894 — Parallelomma )

sellata ( Hackman, 1956 — Chylizosoma ) vittata (Meigen, 1826 — Cordylura )

Megaphthalma Becker, 1894 = Megaphthalmoides Ringdahl, 1936

pallida ( Fallén, 1819 — Cordilura )

unilineata (Zetterstedt, 1838 — Cordilura )

Microprosopa Becker, 1894

= Allomyella Malloch, 1923

albipennis (Zetterstedt, 1838 — Cordilura ) crinipes Ringdahl, 1928

frigida (Holmgren, 1883 — Cordilura ) = varitibia Becker, 1897

haemorrhoidalis (Meigen, 1826 — Cordilura ) heteromyzina (Zetterstedt, 1838 — Scatomyza ) lineata (Zetterstedt, 1838 — Cordilura ) pallidicauda (Zetterstedt, 1838 — Cordilura ) paveli Ozerov et Krivosheina, 2013

portenkoi Stackelberg, 1952

taimyrica Ozerov, 2017

unguiculata (Malloch, 1919 — Allomyia ) zlobini Ozerov, 2009

Micropselapha Becker, 1894

basovi Ozerov, 2010

filiformis (Zetterstedt, 1846 — Cordilura )

Mixocordylura Hendel, 1909

longifacies Hendel, 1909

parva Ozerov et Krivosheina, 2012

Neochirosia Malloch, 1917

anthrax (Schiner, 1864 — Cleigastra )

nigriceps ( Becker, 1894 — Clidogastra ) pechorica Ozerov, 2010

veratri (Hendel, 1925 — Clidogastra )

Norellia Robineau-Desvoidy, 1830

tipularia (Fabricius, 1794 — Musca )

Norellisoma Hendel, 1910

= Norelliosoma , error.

armipes (Meigen, 1826 — Cordylura )

caucasicum (Ozerov, 1993 — Norellia ) flavostriatum Ozerov, 2009

insulare (Ozerov, 1993 — Norellia )

lesgiae ( Becker, 1894 — Norellia )

lituratum (Meigen, 1826 — Cordylura )

= altaica (Ozerov, 2008 — Norellia ) montanopratense (Ozerov, 1993 — Norellia ) oreinum Ozerov, 2010

orientale (Ozerov, 1993 — Norellia )

spinimanum ( Fallén, 1819 — Cordylura ) = septentrionale (Hendel, 1930 — Norelliosoma ) tomkovichi Ozerov, 2010

yolduense (Ozerov, 2008 — Norellia )

Paracosmetopus Hackman, 1956

hellehi Hackman, 1956

Pleurochaetella Vockeroth, 1965

barkalovi Ozerov in Ozerov & Krivosheina, 2012 simplicipes (Becker, 1900 — Cosmetopus )

Pogonota Zetterstedt, 1860

= Lasioscelus Becker, 1894

= Okeniella Hendel, 1907

barbata (Zetterstedt, 1838 — Cordylura )

caudata (Zetterstedt, 1838 — Cordylura ) dasyprocta (Loew, 1864 — Cordylura )

gorodkovi (Ozerov, 2007 — Okeniella )

immunda (Zetterstedt, 1838 — Cordylura ) nigricans (Loew, 1873 — Cordylura )

sahlbergi (Becker, 1900 — Lasioscelus ) stackelbergi ( Gorodkov, 1967 — Okeniella )

Scathophaga Meigen, 1803

= Scopeuma Meigen, 1800

= Conisternum Strobl, 1894

= Coniosternum Becker, 1894

= Scatophaga , unjustified emend.

apicalis (Curtis in Ross, 1835 — Scatophaga ) = arctica (Becker, 1897 — Scatophaga )

= perfecta (Becker, 1907 — Scatophaga )

buryatica Ozerov et Krivosheina, 2019

calida (Haliday in Curtis, 1832 — Scatophaga ) cordylurina (Holmgren, 1883 — Scatomyza ) crinita (Coquillett, 1901 — Scatophaga )

dasythrix ( Becker, 1894 — Scathophaga ) decipiens (Haliday in Curtis, 1832 — Scatophaga ) = fluvialis (Rondani, 1866 — Scatina )

exalata Ozerov, 1996

furcata (Say, 1823 — Pyropa )

incola (Becker, 1900 — Scatophaga )

incompleta Ozerov et Krivosheina, 2020

inquinata (Meigen, 1826 — Scatophaga )

intermedia (Walker, 1849 — Scatophaga )

= hadleyi Ozerov, 2013

karelica Ozerov, 2013

lapponica (Ringdahl, 1920 — Coniosternum )

litorea ( Fallén, 1819 — Scatomyza )

= stuxbergii (Holmgren, 1880 — Scatomyza )

lutaria (Fabricius, 1794 — Musca )

mollis ( Becker, 1894 — Scatophaga )

= sychevskayae Ozerov, 2010

multisetosa (Holmgren, 1883 — Scatomyza )

nigripalpis (Becker, 1907 — Scatophaga )

= orbitalis (Becker, 1915 — Scopeuma )

obscura ( Fallén, 1819 — Cordylura )

obscurinervis (Becker, 1900 — Scatophaga )

pictipennis (Oldenberg, 1923 — Scatophaga )

spurca (Meigen, 1826 — Scatophaga )

= Scatomyza suilla (Fabricius) ; Fallén (1819:3). Misidentification stercoraria (Linnaeus, 1758 — Musca )

= erythrostoma Holmgren, 1883 — Scatomyza )

taeniopa (Rondani, 1867 — Scatophaga )

tinctinervis ( Becker, 1894 — Coniosternum )

varipes (Holmgren, 1883 — Scatomyza )

= septentrionalis (Becker, 1897 — Scatophaga )

yakutica Ozerov, 2017

Scatomyza Fallén, 1810

magnipennis (Portschinsky, 1887 — Scatophaga )

mellipes (Coquillett, 1899 — Scatophaga )

= eoa Ozerov, 2007b ( Scathophaga )

scybalaria (Linnaeus, 1758 — Musca )

Spaziphora Rondani, 1856

hydromyzina ( Fallén, 1819 — Cordylura )

tomkovichi Ozerov, 2012

Staegeria Rondani, 1856

kunzei (Zetterstedt, 1821 — Cordylura )

Trichopalpus Rondani, 1856

= Chaetosa Coquillett, 1898

= Paramicroprosopa Ringdahl, 1936

= Huckettia Vockeroth, 1995

= Tricopalpus , error fraternus (Meigen, 1826)

lacteipennis (Ringdahl, 1920 — Microprosopa )

nearcticus (Vockeroth, 1995 — Huckettia )

nigribasis (Curran, 1927 — Tricopalpus )

obscurellus (Zetterstedt, 1846 — Cordylura )

punctipes (Meigen, 1826 — Cordylura )

Acknowledgements. We are grateful to Dmitry I. Gavryushin (Moscow) for the use of images of living Cleigastra apicalis ( Fig. 105 ), C. planiceps ( Fig. 108), Cordilura albiceps ( Fig. 109), C. picipes ( Fig. 110), Hydromyza livens ( Fig. 115 ), Leptopa filiformis ( Fig. 116 ), Megaphthalma pallida ( Fig. 117 ), Norellisoma spinimanum ( Fig. 120 ), Scathophaga furcata ( Fig. 122 ), S. obscura ( Fig. 123 ), S. stercoraria ( Fig. 124 ), Trichopalpus fraternus ( Fig. 127 ) and Trichopalpus punctipes ( Fig. 128 ).

The investigation was fulfilled within the state project of the Institute of the Ecology and Evolution RAS (M.G. Krivosheina) and the State project of Lomonosov Moscow State University No 121032300105-0 (A.L. Ozerov) .

Figures

Figs 19–21. Head of Scathophagidae, lateral view (19, 20) and dorsolateral view (21): 19 — Spaziphora hydromyzina (Fallén); 20 — Trichopalpus nearcticus (Vockeroth); 21 — Hydromyza livens (Fabricius). Рис. 19–21. Голова Scathophagidae, сбоку (19, 20) и дорсолатерально (21): 19 — Spaziphora hydromyzina (Fallén); 20 — Trichopalpus nearcticus (Vockeroth); 21 — Hydromyza livens (Fabricius).

Figs 22–27. Fragments of thorax of Scathophagidae: 22 — Hydromyza livens (Fabricius), anepisternum; 23 — Trichopalpus fraternus (Meigen), anepisternum; 24 — Scathophaga litorea (Fallén), anepisternum; 25 — Cleigastra flavipes (Fallén), anepisternum and katepisternum; 26 — Cleigastra arctica (Becker), anepisternum and katepisternum; 27 — Trichopalpus punctipes (Meigen), anepisternum and katepisternum. Abbreviations: a kepst s — anterior katepisternal seta; anepst — anepisternum; kepst — katepisternum; lp kepst s — lower posterior katepisternal seta; prepst — proepisternum; up kepst s — upper posterior katepisternal seta. 24 — after Ozerov & Krivosheina, 2021, fig. 6; 25 — after Ozerov & Krivosheina, 2015, fig. 1. Рис. 22–27. Фрагменты груди Scathophagidae: 22 — Hydromyza livens (Fabricius), анЭпистерн; 23 — Trichopalpus fraternus (Meigen), анЭпистерн; 24 — Scathophaga litorea (Fallén), анЭпистерн; 25 — Cleigastra flavipes (Fallén), анЭпистерн и катЭпистерн; 26 — Cleigastra arctica (Becker), анЭпистерн и катЭпистерн; 27 — Trichopalpus punctipes (Meigen), анЭпистерн и катЭпистерн. ОбоЗначениЯ: a kepst s — переднЯЯ катЭпистернальнаЯ Щетинка; anepst — анЭпистернум; kepst — катЭпистернум; lp kepst s — нижнЯЯ ЗаднЯЯ катЭпистернальнаЯ Щетинка; prepst — проЭпистернум; up kepst s — верхнЯЯ ЗаднЯЯ катЭпистернальнаЯ Щетинка. 24 — по Ozerov & Krivosheina, 2021, fig. 6; 25 — по Ozerov & Krivosheina, 2015, fig. 1.

Figs 124–128. Habitus of Scathophagidae: 124 — Scathophaga stercoraria (Linnaeus); 125 — Spaziphora hydromyzina (Fallén); 126 — Staegeria kunzei (Zetterstedt); 127 — Trichopalpus fraternus (Meigen); 128 — Trichopalpus punctipes (Meigen). 124 — after Ozerov & Freidberg, 2011, fig. 24; 127, 128 — after Ozerov, 2019c, figs 1, 2. Рис. 124–128. ОбЩий вид Scathophagidae: 124 — Scathophaga stercoraria (Linnaeus); 125 — Spaziphora hydromyzina (Fallén); 126 — Staegeria kunzei (Zetterstedt); 127 — Trichopalpus fraternus (Meigen); 128 — Trichopalpus punctipes (Meigen). 124 — по Ozerov & Freidberg, 2011, fig. 24; 127, 128 — по Ozerov, 2019c, figs 1, 2.

Figs 100–105. Habitus of Scathophagidae: 100 — Acanthocnema altaica Ozerov & Krivosheina; 101 — Acerocnema breviseta (Zetterstedt); 102 — Acerocnema nitens Ozerov & Krivosheina; 103 — Acerocnema richterae Ozerov; 104 — Bostrichopyga crassipes (Zetterstedt); 105 — Cleigastra apicalis (Meigen). 100 — after Ozerov & Krivosheina, 2018a, fig. 1; 102 — after Ozerov & Krivosheina, 2018b, fig. 6; 103 — after Ozerov, 2015, fig. 1; 104 — after Ozerov & Krivosheina, 2014a, fig. 1; 105 — after Ozerov, 2013, fig. 19. Рис. 100–105. ОбЩий вид Scathophagidae: 100 — Acanthocnema altaica Ozerov & Krivosheina; 101 — Acerocnema breviseta (Zetterstedt); 102 — Acerocnema nitens Ozerov & Krivosheina; 103 — Acerocnema richterae Ozerov; 104 — Bostrichopyga crassipes (Zetterstedt); 105 — Cleigastra apicalis (Meigen). 100 — по Ozerov & Krivosheina, 2018a, fig. 1; 102 — по Ozerov & Krivosheina, 2018b, fig. 6; 103 — по Ozerov, 2015, fig. 1; 104 — по Ozerov & Krivosheina, 2014a, fig. 1; 105 — по Ozerov, 2013, fig. 19.

Figs 112–117. Habitus of Scathophagidae: 112 — Neochirosia veratri (Hendel); 113 — Ernoneura argus (Zetterstedt); 114 — Gimnomera tarsea (Fallén); 115 — Hydromyza livens (Fabricius); 116 — Leptopa filiformis Zetterstedt; 117 — Megaphthalma pallida (Fallén). 114 — after Ozerov, 2019a, fig.1. Рис. 112–117. ОбЩий вид Scathophagidae: 112 — Neochirosia veratri (Hendel); 113 — Ernoneura argus (Zetterstedt); 114 — Gimnomera tarsea (Fallén); 115 — Hydromyza livens (Fabricius); 116 — Leptopa filiformis Zetterstedt; 117 — Megaphthalma pallida (Fallén). 114 — по Ozerov, 2019a, fig.1.

Figs 118–123. Habitus of Scathophagidae: 118— Microprosopa frigida (Holmgren); 119 — Microprosopa pallidicauda (Zetterstedt); 120 — Norellisoma spinimanum (Fallén); 121 — Paracosmetopus helleni Hackman; 122 — Scathophaga furcata (Say); 123 — Scathophaga obscura (Fallén). 118, 119 — after Ozerov, 2017, figs 2, 1; 120 — after Ozerov, 2010a, fig 1; 121 — after Ozerov & Krivosheina, 2014a, fig. 8; 122, 123 — after Ozerov & Krivosheina, 2021, figs 3, 1. Рис. 118–123. ОбЩий вид Scathophagidae: 118— Microprosopa frigida (Holmgren); 119 — Microprosopa pallidicauda (Zetterstedt); 120 — Norellisoma spinimanum (Fallén); 121 — Paracosmetopus helleni Hackman; 122 — Scathophaga furcata (Say); 123 — Scathophaga obscura (Fallén). 118, 119 — по Ozerov, 2017, figs 2, 1; 120 — по Ozerov, 2010a, fig 1; 121 — по Ozerov & Krivosheina, 2014a, fig. 8; 122, 123 — по Ozerov & Krivosheina, 2021, figs 3, 1.