Angustopila werneri Pall-Gergely & Hunyadi, 2023

Pall-Gergely, Barna, Hunyadi, Andras, Vermeulen, Jaap J., Grego, Jozef, Sutcharit, Chirasak, Reischuetz, Alexander, Dumrongrojwattana, Pongrat, Botta-Dukat, Zoltan, Oerstan, Aydin, Fekete, Judit & Jochum, Adrienne, 2023, Five times over: 42 new Angustopila species highlight Southeast Asia's rich biodiversity (Gastropoda, Stylommatophora, Hypselostomatidae), ZooKeys 1147, pp. 1-177 : 1

Angustopila werneri Pall-Gergely & Hunyadi sp. nov.

Figs 25 , 26

Type material.

Holotype: Vietnam • 1 empty shell (H: 1.09 mm, D: 1.01 mm); Lạng Sơn Province, Bắc Sơn District, Quinh Son, southern edge of the village, left side of road no. 241 (locality code: 2020/49); 21°53.41'N, 106°20.94'E; 400 m a.s.l.; 18 Feb. 2020; A. Hunyadi leg.; HNHM 105280.

Paratypes: Vietnam • 10 adult shells; same data as for holotype; coll. HA.

Additional material.

Vietnam • 4 juvenile shells; same data as holotype; coll. HA • 7 shells; Lạng Sơn Province, H ữu Lũng District, H ữu Liên, 1400 m west from Đȏng Lâm along road no. 241 (locality code: 2020/53); 21°41.91'N, 106°21.77'E; 210 m a.s.l.; 19 Feb. 2020; A. Hunyadi leg.; coll. HA .

Diagnosis.

A medium- to large-sized, concave-conical Angustopila species with a comparatively small, toothless aperture and a wide umbilicus.

Description.

Shell medium to large-sized for the genus, slightly higher than wide or slightly wider than high; concave-conical; last whorl widest from standard apertural view; protoconch consists of 1.25 whorls, with strong spiral striae; teleoconch with some fine, weak, radial growth lines and much stronger, elevated, dense, mostly regularly spaced spiral striae (ca. 18-20 on body whorl in standard apertural view); whorls 4.25, rounded, or body whorl slightly shouldered; aperture strongly oblique to shell axis from lateral view; umbilicus wide; aperture relatively small, subcircular, irregular in shape, parietal part straight or convex; peristome slightly expanded, not reflected; parietal callus considerably detached from penultimate whorl; aperture toothless.

Measurements (in mm).

H = 0.94-1.09, D = 0.92-1.01, H/D*100 = 97-110.4 (n = 8), RUD = 29.7-34.7 (n = 4).

Differential diagnosis.

The most similar species in terms of shell and aperture shape is A. elevata , which is much smaller, has a more conical (rather than concave-conical) shell resulting in a narrower umbilicus, and less prominent radial sculpture. See also under A. halongensis sp. nov.

Etymology.

The new subspecies is dedicated to and named after Ervin Werner (1936-2021), the prominent botanist and biology teacher in secondary school of the first author.

Distribution.

The two known populations are located ca. 20 kilometres from each other in Lạng Sơn Province, northern Vietnam (Fig. 27 ).

Remarks.

Shells from locality 2020/53 have a comparatively smaller aperture than those of the typical shells, but no other differences were found.

Figures

Figure 25. Angustopila werneri Pall-Gergely & Hunyadi, sp. nov., (holotype, HNHM 105280). Apertural (A), lateral (B), apical (C), and ventral (D) sides of the shell; sculpture on the protoconch (E), ventral (F) and frontal (G) surface of the body whorl.

Figure 26. Angustopila werneri Pall-Gergely & Hunyadi, sp. nov. 2020 / 53 (paratypes) A specimen 2, B-H specimen 1. Apertural (A, B), lateral (C), ventral (D) and apical (E) sides of the shell; sculpture on the protoconch (F), ventral (G) and frontal (H) surface of the body whorl.

Figure 27. Distribution of Angustopila species.