Scambus irapuatoensis Khalaim & Ruíz-Cancino, 2022

Khalaim, Andrey I. & Ruíz-Cancino, Enrique, 2022, Darwin wasps of the subfamily Pimplinae (Hymenoptera: Ichneumonidae) of Mexico: Ephialtes genus-group, Zootaxa 5169 (3), pp. 201-251 : 237

8. Scambus irapuatoensis Khalaim & Ruíz-Cancino , sp. nov.

( Figs 40 , 59 )

Material examined. Holotype female ( EMEC), Mexico, Guanajuato, 18 mi. W of Irapuato , 24.vii.1954, coll. J.W. MacSwain & E.I. Schlinger.

Paratypes. MEXICO. Coahuila: 3 ♀ (2 in AEIC, 1 in ZISP) 6 mi. W of Saltillo , 5200 ft. (= 1585 m), 14– 20.vii.1972, coll. B. & C. Dasch. Nuevo León: 1 ♀ (TAMU) 12.4 mi. NE of Dr. Arroyo , 7.vii.1986, coll. Schaffner & Kovarik. Michoacán: 1 ♀ ( EMEC) 10 km W of Zitácuaro , 11.vii.1951, coll. P.D. Hurd. México: 1 ♀ (UAT) Jilotepec , 14.vii.1992, coll. A.J. Guzmán Larralde. 1 ♀ ( FSCA) ca. [San Gregorio] Atlapulco , 3000 m, 16.vi.1974, coll. C. Porter. Puebla: 1 ♀ (UAT) Cholula , pastos, en cultivo, 15.v.1992, coll. A.J. Guzmán Larralde. 8 ♀ ( AEIC) Cholula , 8–11.xi.1972, coll. B. & C. Dasch.

Description. Female ( Fig. 40 ). Fore wing length 7.6 mm.

Body length 9.0 mm. Head in dorsal view almost twice as broad as long, strongly rounded behind eyes in dorsal view. Mandible weakly tapered towards apex, with upper tooth somewhat longer and much broader than the lower. Clypeus with lower margin medially strongly concave. Malar space about 0.3× as long as basal mandibular width. Face distinctly transverse, 1.45× as broad as medially high, weakly convex. Lateral ocellus separated from eye by 1.3× its own maximum diameter. Head polished with fine and very sparse setiferous punctures.

Mesoscutum polished, finely punctate and moderately densely pubescent. Epomia weak, curved. Notaulus deeply impressed, long, reaching centre of mesoscutum. Submetapleural carina distinct, complete; metapleuron rather short. Propodeum mediodorsally polished, laterally with shallow punctures; lateromedian longitudinal carinae represented by short ridges in anterior 0.2 of propodeum. Pleural carina present, fine but quite distinct behind propodeal spiracle. Legs slender, hind coxa 1.45× as long as maximally broad.

Fore wing with 1cu-a (nervulus) opposite to M&RS (basal vein). Hind wing with nervellus distinctly intercepted in posterior 0.4, distal section of CU weakly pigmented.

First tergite 0.95×, second tergite 0.65× and third tergite 0.6× as long as posteriorly (maximally) broad. First tergite in profile with dorsal margin straight in basal 0.7, abruptly angled and slightly convex posteriorly; lateromedian carinae distinct in basal 0.7 of tergite; lateral longitudinal carina more or less complete, sometimes partly obliterated. Tergites 2–5 with distinct and dense punctures, with smooth or finely striate impunctate bands on posterior 0.2. Tergite 5 weakly and tergite 6 strongly emarginated posteromedially. Ovipositor straight, robust, moderately compressed, apically with weak nodus and about 8–9 oblique teeth on lower valve; most proximal teeth strongly oblique. Sheath 2.3× as long as hind tibia.

Head black, clypeus and base of mandible dark reddish brown. Antenna black with slight brownish tinge. Mesosoma black with only hind corner of pronotum narrowly white ( Fig. 59 ). Tegula white. Pterostigma pale yellow. Wings yellowish. Legs uniformly reddish orange, hind tarsomeres sometimes slightly infuscate apically ( Fig. 59 ). Tergite 1 black. Following metasomal tergites reddish brown laterally to black medially, with posterior 0.2 black (black band more or less reduced medially on tergites 5 and 6).

Male. Unknown.

Distribution. Mexico (Coahuila, Guanajuato, Nuevo León, Michoacán, Mexico, Puebla).

Comparison. Runs to S. canadensis Walley in couplet 10 of the key to the Nearctic species by Townes & Townes (1960: 39), but differs from this species by the pale yellow pterostigma (brown in S. canadensis ), uniformly reddish orange legs (with extensive blackish marking, see Fig. 320k in Townes & Townes 1960: 616), white hind corner of pronotum, and propodeum with very short lateromedian longitudinal carinae.

Figures

FIGURES 37–40. Habitus, lateral, ♀: 37—Acropimpla tequila sp. nov. (holotype); 38—Scambus aplopappi; 39—S. longicoxa sp. nov. (paratype); 40—S. irapuatoensis sp. nov. (paratype).

FIGURES 59–62. Scambus irapuatoensis sp. nov., paratype, ♀: 59—head, mesosoma and base of metasoma, lateral. Scambus longicoxa sp. nov., holotype, ♀: 60—habitus (without wing apices), lateral; 61—hind coxa, lateral; 62—apex of ovipositor, lateral.