Sarika benoiti (Crosse & Fischer, 1863)

Pholyotha, Arthit, Sutcharit, Chirasak, Tongkerd, Piyoros & Panha, Somsak, 2020, Integrative taxonomic revision of the land snail genus Sarika Godwin-Austen, 1907 in Thailand, with descriptions of nine new species (Eupulmonata, Ariophantidae), ZooKeys 976, pp. 1-100 : 1

Sarika benoiti (Crosse & Fischer, 1863) Fig. 50F

Zonites benoiti Crosse & Fischer, 1863: 346, pl. 14, fig. 4. Type locality: "in loco Fuyen-Moth dicto, Cochinchine" [Phu Yen Province, Vietnam].

Helix benoiti : Pfeiffer 1868: 99; Morelet 1875: 248.

Nanina (Macrochlamys) benoiti : Tryon 1886: 90, pl. 30, fig. 57, 58.

Ariophanta ( Macrochlamys ?) benoiti : Fischer 1891: 20.

Macrochlamys benoiti : Ancey 1898: 128, pl. 9, Fig. b; Saurin 1953: 113; Schileyko 2011: 30, 31.

Sarika benoiti : Inkhavilay et al. 2019: 81, fig. 38c; Pholyotha et al. 2020a: 7.

Type material.

No type material could be located in the MNHN collections. The illustration from the original description is reproduced here (Fig. 50F ).

Diagnosis.

Shell depressed, medium size (shell width up to 16.0 mm, shell height up to 9.0 mm), and thin. Shell surface smooth and shining; shell colour brownish. Whorls 6, increasing regularly; body whorl well rounded. Spire elevated; suture impressed. Aperture crescent-shaped and obliquely opened. Peristome simple. Columellar margin simple. Umbilicus narrowly opened (Fig. 50F ).

Remarks.

This species was described based on a specimen from south-central Vietnam ( Crosse and Fischer 1863). Up to now, Sarika benoiti has been recorded throughout the Indochina countries ( Ancey 1898, Fischer and Dautzenberg 1904, Saurin 1953, Schileyko 2011, Inkhavilay et al. 2019). However, these records remain equivocal and tentative because of unavailable distinguished genital data. In Thailand, Sarika benoiti was recorded from eastern Thailand based on specimens from the Pavie collection ( Fischer and Dautzenberg 1904).

In this study, the specimens from the eastern part of Thailand agreed well with the syntype of S. bocourti (Fig. 34A ) where its type locality is in Battambang, Cambodia. Based on shell morphology, S. benoiti (Fig. 50F ) differs slightly from S. bocourti in having a smaller sized and well-rounded body whorl, whereas S. bocourti has a larger sized shell (shell width up to 33 mm) and obtusely angulated body whorl.

Figures

Figure 34. Shells of Group II: Sarika hainesi group. A-C Sarika hainesi A syntype NHMUK 20200290 and B, C specimen CUMZ 7237 D-F S. bocourti D syntype NHMUK 1893.2.4.1076 - 1077 E specimen CUMZ 7590, and F specimen CUMZ 7592.

Figure 50. Shells of Sarika spp. A, B Sarika gratesi sp. nov. A holotype CUMZ 7275 and B paratype CUMZ 7276 C, D S. pumicata syntype NHMUK 1893.2.4.980 - 981 E S. ochtogyra holotype SMF 227103 / 1 F S. benoiti modified from Crosse and Fischer (1863) G S. rex modified from Preston (1909).