Clitellariinae, Brauer, 1882

Winterton, Shaun L., 2023, Wasp-mimicking soldier flies of the Australian Region: revision of Ampsalis Walker, Elissoma White and Lagenosoma Brauer (Stratiomyidae: Clitellariinae), Zootaxa 5246 (1), pp. 1-63 : 9-10

Key to Clitellariinae genera of the Oriental and Australian regions

(including the Indo-Malayan transition zone)

1. Notopleural spine present............................................................................... 2.

- Notopleural spine absent............................................................................... 4.

2. Apical stylus of antenna with dense and elongate setae (widely distributed from Australia to China; a single Afrotropical species)............................................................................. Nigritomyia Bigot, 1877 .

- Apical stylus of antenna bare or pubescent, without elongate setae.............................................. 3.

3. Abdomen broadly rounded ( Indonesia; widely distributed throughout Oriental and Palaearctic regions).................................................................................................. Clitellaria Meigen, 1800 .

- Abdomen elliptical ( Indonesia)...................................................... Anoamyia Lindner, 1935 .

4. Head relatively flattened dorsoventrally in profile, longer than high............................................. 5.

- Head broadly rounded, or taller and long in profile........................................................... 6.

5. Face flat, not produced beyond eye margin in lateral view; antenna short and aristate; wing extends well beyond end of abdomen ( New Zealand)................................................................. Dysbiota Lindner, 1958 .

- Face produced anteriorly around base of antenna with a triangular ventral tooth-like projection; antennae elongate, nearly all flagellomeres of equal size and not aristate; wing not extending far beyond end of abdomen ( Indonesia, Papua New Guinea)........................................................................... Caenocephaloides Strand, 1898 .

6. Scutellum without spines............................................................................... 7.

- Scutellum with marginal spines present.................................................................... 9.

7. Abdomen round; antenna relatively short, similar to head length; flagellum tapered apically; body mostly bright orange ( Indonesia, Papua New Guinea, India).......................................................... Ruba Walker, 1859 .

- Abdomen ovoid to elongate; antenna usually much longer than head, flagellum uniform in width along length; body colour variable............................................................................................. 8.

8. Abdomen ovoid to petiolate (more so in male), anterior segments (1–3) much narrower than posterior segments (4–8) (e.g., Fig. 33 ); face frequently protruding and observable in lateral view ( Australia).................... Lagenosoma Brauer, 1882 .

- Abdomen ovoid, never petiolate ( Fig. 34D ); face not protruding and barely observable in lateral view ( China, India, Indonesia).......................................................................... Eudmeta Wiedemann, 1830 .

9. Wing with cross-vein r–m absent; small, predominantly black species with wings folded over abdomen ( Australia, Papua New Guinea).......................................................................... Octarthria Brauer, 1882 .

- Wing with cross-vein r–m present; body size variable and wings usually straight.................................. 10.

10. Flagellum with flagellomeres 4–7 short and strongly tapered, flagellomere 8 very narrow and elongate with terminal style; individuals with black purplish colouration ( Australia)..................................... Geranopus White, 1916 .

- Flagellum elongate with flagellomeres uniformly cylindrical, variable in length and shape, lacking hair-like terminal style..................................................................................................... 11.

11. Scutellar spines greatly elongate (longer than scutellum length) and strongly divergent in orientation ( Indonesia, Malaysia, Thailand, India, Laos)......................................................... Campeprosopa Macquart, 1850 .

- Scutellar spines shorter than scutellum length, sometimes minute, parallel or only moderately divergent in orientation… 12 ...

12. Wing vein M 4 emerging before discal cell ( China, Indonesia, Malaysia, India, Papua New Guinea; also found in Neotropical and Nearctic regions).......................................................... Cyphomyia Wiedemann, 1819 .

- Wing vein M 4 emerging from discal cell.................................................................. 13.

13. Eyes pilose......................................................................................... 14.

- Eyes not pilose...................................................................................... 15.

14. Body black and yellow with extensive yellow setal pile, excellent Vespidae mimics; antenna much longer than head; all flagellomeres similar in shape and width, flagellomeres 1–7 with circular sensory pits ( Australia)... Syndipnomyia Kertész, 1921 .

- Body dark in colouration; antenna similar length to head; basal flagellomeres much thicker than distal flagellomeres, flagellomeres 1–3 with circular sensory pits (widely distributed except Australasia)................. Adoxomyia Kertész, 1907 .

15. Face below antennae flat or rounded, without ‘nose’-like process directed anteroventrally (e.g., Fig. 25E–N ) ( Australia)....................................................................................... Elissoma White, 1916 .

- Face below antennae with distinct ‘nose’-like anteroventrally directed process ( Indonesia, Australia) (e.g., Fig. 25A–D )........................................................................... Ampsalis Walker, 1859 sensu stricto.

Figures

FIGURE 25. Heads of Ampsalis and Elissoma. A. Ampsalis acta sp. n., male; B. Ampsalis acta sp. n., female; C. Ampsalis wallacei sp. n., male; D. Ampsalis wallacei sp. n., female; E. Elissoma australis sp. n., female; F. Elissoma gilva sp. n., female; G. Elissoma danielsi sp. n., male; H. Elissoma danielsi sp. n., female; I. Elissoma ichneumonoides sp. n., female; J. Elissoma hespera sp. n., male; K. Elissoma brunnea Hardy, male; L. Elissoma brunnea Hardy, female; M. Elissoma minuta sp. n., male; N. Elissoma lauta White, male.

FIGURE 33. Lagenosoma abdomens. A. Lagenosoma danielsorum sp. n., male; B. Lagenosoma danielsorum sp. n., female; C. Lagenosoma dimorphum sp. n., male; D. Lagenosoma dimorphum sp. n., female; E. Lagenosoma dispar Brauer, male; F. Lagenosoma dispar Brauer, female; G. Lagenosoma pictum Brauer, male; H. Lagenosoma pictum Brauer, female; I. Lagenosoma propinquum Brauer, male; J. Lagenosoma propinquum Brauer, female; K. Lagenosoma varigatum sp. n., male; L. Lagenosoma varigatum sp. n., female.

FIGURE 34. Abdomens. A. Lagenosoma eklemmum sp. n., female; B. Lagenosoma propius sp. n., male; C. Eudmeta marginata (Fabricius), male; D. Campeprosopa longispina (Brunetti, 1913), female.