Deuterothynnus, Brown, 2005

Brown, G. R., 2005, A revision of Tachyphron Brown and description of two new genera within the Ariphron group (Hymenoptera: Tiphiidae), Journal of Natural History 39 (2), pp. 197-239 : 203-205

Deuterothynnus gen. nov.

Type species: Deuterothynnus fulvisetatus sp. n.

Etymology. The generic name is derived from the Greek word deuteros meaning second, and is a reference to both a second large apical spine on the hypopygium and a secondary smaller preapical spine.

Diagnosis. Male: hypopygium deeply emarginate medially between two long spines, and a smaller preapical spine on the outer margin of these long spines.

Description. Male: similar to Tachyphron , but differs by having the hypopygium deeply emarginate medially between two long spines, a smaller preapical spine on the outer margin of these long spines ( Figures 13–16 ) and the genitalia ( Figures 5–12 ) with (1) the basiparameres long and (collectively) narrowly triangular or narrowly subparallel with the lateral margins straight over most of their length; (2) parameres (viewed laterally) usually broad over most of their length; and (3) the aedeagus differentiated into a small basal hoodlike structure without ventral lobes and apically into a long and often convoluted filament.

Remarks. Four species, D. fulvicentratus , D. fulvisetatus , D. insularis and D. parallelus , from New Guinea (including Irian Jaya) or the adjacent island of Misool are known ( Figure 59 ). All are recorded from a single locality and from at most two specimens.

The only known female specimen, the holotype of Deuterothynnus insularis (Turner) comb. nov., apparently has been lost, and the original description is insufficient to provide a diagnosis.

Key to males of Deuterothynnus (only the female of D. insularis is known but the holotype could not be located)

1.

– Metasomal segments 1 and 2 orange, remainder of metasoma black Metasoma concolorous, dark brown to black............... 2 3 2.

–

Genitalia with basiparameres (collectively) with lateral margins subparallel (Figure

12) and parameres with dorsal margin gently curved over most of length ( Figure 11 ); setae white............... fulvicentratus sp. n. Genitalia with basiparameres (collectively) with lateral margins subtriangular ( Figure 10 ) and parameres with dorsal margin sinusoidal on basal half ( Figure 9 ); setae orange................ fulvisetatus sp. n. 3. Propodeum shallowly punctate; genitalia with basiparameres (collectively) with –

lateral margins subtriangular ( Figure 8 ) and parameres with ventral margin straight with apex straight and narrow ( Figure 7 )........ insularis Turner Propodeum closely to rugosely punctate; genitalia with basiparameres (collectively) with lateral margins subparallel ( Figure 6 ) and parameres with ventral margin curved with apex curved and broad ( Figure 5 )....... parallelus sp. n.

Figures

Figure 59. Distribution of Deuterothynnus spp. and Heligmothynnus spp. (note the coincident collecting of D. fulvicentratus with H. microspinus, and D. fulvisetatus with D. parallelus).

Figures 13–16. Deuterothynnus spp., male hypopygia. (13) D. fulvicentratus. (14) D. fulvisetatus. (15) D. parallelus. (16) D. insularis. Scale bar: 0.25 mm.

Figures 5–12. Deuterothynnus spp., male genitalia, lateral and dorsal views. (5, 6) D. parallelus. (7, 8) D. insularis. (9, 10) D. fulvisetatus. (11, 12) D. fulvicentratus. a, Aedeagus; b, right basiparameres; c right cuspis; p, right parameres; r, basal ring. Scale bars: 0.25 mm.