Coptocheilus sumatranus Dohrn, 1881

Jirapatrasilp, Parin, Sutcharit, Chirasak & Panha, Somsak, 2022, Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia, ZooKeys 1119, pp. 1-115 : 1

Coptocheilus sumatranus Dohrn, 1881

Fig. 5D-F

Coptocheilus sumatranus Dohrn, 1881: 65. Type locality: Sumatra, Singalang [Mount Singgalang, West Sumatra].

Megalomastoma sectilabrum [in part]- Stoliczka 1872: 268, pl. 10, fig. 13, Penang hill [Penang Island, Penang State, Malaysia]. Crosse 1879: 339, Perak [Malaysia]. de Morgan 1885: 412, 413.

Megalomastoma (Coptocheilus) sectilabrum [in part]- Nevill 1878: 297.

Megalomastoma (Coptochilus) sectilabrum [in part]- von Martens 1886: 161. von Möllendorff 1887 [1886]: 314; Larut [Bukit Larut, Perak State, Malaysia].

Coptochilus sectilabrum [non Gould]- von Möllendorff 1891: 346.

Schistoloma sectilabrum [in part]- Kobelt 1902: 280. Gude 1921: 170, 171.

Coptocheilus perakensis Fulton, 1903: 102, pl. 9, fig. 3. Type locality: Perak.

Schistoloma perakense - Laidlaw 1928: 33.

Schistoloma sectilabrum [non Gould]- Sykes 1903: 197, Ulu Selama, Perak. Laidlaw 1928: 33, Ulu Selama, Perak; Lampan Patalung [Phatthalung Province, Thailand]. Foon et al. 2017: 41, fig. 16b, Perak, forested slope behind the village at Gunung Pondok.

Schistoloma sumatranum - Kobelt 1902: 281. van Benthem Jutting 1949: 55, 56, Kuala Legap, Plus Valley, Perak; Maxwell’s Hill, Perak; Gunong Kledang, Perak; Taiping Perak; Dusun Tua, Selangor [Malaysia]. Davison 1995: 236, Sungai Halong and Sungai Emban, Temengor Forest Reserve, Perak, Malaysia. Chan 1998a: 4, Ipoh, Perak. Maassen 2001: 43, 44. Páll-Gergely et al. 2019: 327.

Schistoloma perakensis - Berry 1963: pl. 6, fig. 29.

Type material examined.

Syntype of Coptocheilus perakensis NHMUK 1903.11.20.33 (1 shell; Fig. 5D ) from Perak.

Other material examined.

SMF 262529/1 " Schistoloma siamensis Brandt" (1 shell; Fig. 5E ) from Thailand: an den Tanto-Fällen bei Ban Nong Star; Yala Provinz [Than To Waterfall Forest Park, Bannang Sata District, Yala Province, Thailand]. NHMUK 1986.4.19.14 " Coptocheilus sectilabrum var." (1 shell; Fig. 5F ) from Larut near Perak.

Diagnosis.

Shell elongate conical without any periumbilical keel. Apertural round without any indentation.

Differential diagnosis.

Coptocheilus sumatranus is different from C. sectilabris in having a round aperture without any indentation.

Distribution.

Peninsular Malaysia, Sumatra Island, and southern Thailand ( Laidlaw 1928; van Benthem Jutting 1949; Foon et al. 2017).

Remarks.

No material of this species was found during this survey. Although C. sumatranus only differs from C. sectilabris by an absence of an indentation in the peristome ( van Benthem Jutting 1949), we do not synonymise C. sumatranus with C. sectilabris because of the lack of DNA data and that there are too few specimens to verify whether specimens collected from the same localities of C. sectilabris eventually lack an angular indentation in the peristome. Coptocheilus perakensis was retrieved as a junior subjective synonym of C. sumatranus because there are no distinct differences in shell form and size between them ( C. sumatranus : shell height 19-24 mm, diameter 8-9 mm; C. perakensis : shell height 23 mm, diameter including peristome 11 mm; van Benthem Jutting 1949).

The name " Schistoloma siamensis Brandt" given to two samples (SMF 262529 = holotype" and SMF 262530 = “paratypes”) from Than To Waterfall Forest Park, Bannang Sata District, Yala Province, Thailand was never published and so is not available. These specimens are larger, more elongated, and have a darker shell colour but the other diagnostic characters conform to those found in the syntype of ' C. perakensis '. Thus, Brandt’s specimens are herein identified as C. sumatranus .

Figures

Figure 5. A-C Coptocheilus sectilabris: A lectotype MCZ 169361 from Tavoy B specimen SMF 109813 from Tavoy, and C specimen CUMZ OLM- 0111 from Kaeng Krachan, Phetchaburi D-F Coptocheilus sumatranus: D syntype of Coptocheilus perakensis NHMUK 1903.11.20.33 from Perak E specimen SMF 262529 / 1 " Schistoloma siamensis Brandt " from Thailand: an den Tanto-Faellen bei Ban Nong Star; Yala Provinz, and F specimen NHMUK 1986.4.19.14 " Coptocheilus sectilabrum var. " from Larut near Perak.