Scaphodius megalognathus, Schöller, 2009

Schöller, M., 2009, Revision of the New Caledonian genus Scaphodius CHAPUIS (Coleoptera: Chrysomelidae: Cryptocephalinae), Linzer biologische Beiträge 41 (1), pp. 767-801 : 782-783

publication ID

https://doi.org/ 10.5281/zenodo.5276273

persistent identifier

https://treatment.plazi.org/id/7425174F-FFE7-F75D-FFD9-17FCAB7AF9E4

treatment provided by

Carolina

scientific name

Scaphodius megalognathus
status

sp. nov.

Scaphodius megalognathus nov.sp.

H o l o t y p e Male (MNHN): Nouvelle-Calédonie, Ouégoa/Mandjélia, Lat. S 20.39683 Long. E 164.53218, Alt. 787 m [orange with black border] / Collecté sur: Divers COL/X03/05, Le 07 et 08/02/05, S. Cazeres & C. Mille [orange with black border] / (my label) Holotypus Scaphodius megalognathus nov.sp. des. Matthias Schöller [red] /.

S i x p a r a t y p e s: 1♀ (MNHN), 2♀♀ (CRSRFP), 1♀ (CRIRD), 1♀ (ZMHB), 1♀ (MESC): Nouvelle-Calédonie, Ouégoa/Mandjélia, Lat. S 20.39683 Long. E 164.53218, Alt. 787 m [orange with black border] / Collecté sur: Divers COL/X03/05, Le 07 et 08/02/05, S. Cazeres & C. Mille [orange with black border] / (all with my label) Paratypus Scaphodius megalognathus nov.sp. des. Matthias Schöller [red] /.

D i a g n o s i s.Alargeblackishbrown,mattspecieswith clypeus, lateral marginsof pronotum, a pair of transverse markings at base of pronotum and a spot on elytron yellow, disc of pronotum glabrous, elytra setose, no punctures in rows, interstices smooth, glabrous, setae short and restricted to furrows and apex of elytra, apex of male clypeus extended, and labrum rectangular, apically straight.

T y p e l o c a l i t y. Nouvelle-Calédonie, Ouégoa/Mandjélia, S 20°39’, E 164°53’, 787 m.

D e s c r i p t i o n o f h o l o t y p e (male) ( Figs 16, 17 View Figs 14-23 , 29 View Figs 24-32 , 49 View Figs 49-52 )

H a b i t u s. Black, matt, with clypeus, lateral margins of pronotum, a pair of transverse markings at base of pronotum and a spot on elytron yellow, body large, shape cylindrical, truncate ( Fig. 49 View Figs 49-52 ), pronotum wider than base of elytra, pronotum glabrous, elytral striae with short white setae; size [mm]: length 5.5, width of elytra at humeri 3.1, length of elytra 3.0; length of pronotum 1.8, maximum width 3.3.

H e a d. Large, hypognathous, sunk into thorax up to hind margin of eyes, puncturation moderately dense and coarse, with an impunctate longitudinal stripe at middle, eyes almost round and relatively small, distance between upper lobes of eyes 2.2 times eye length; antennae inserted near lower margin of eyes, antennae black, scape and pedicel yellowish brown, antennae with all segments elongate, first segment very long, nearly as long as following three segments combined, except for short pedicel, antennomeres expanded from 6 to 10 (10-11 of left antenna missing in holotype), interantennal space seven times diameter of antennal socket; clypeus broad, yellow, basal margin semicircular concave, covered with short white setae, apically with a pair of long setae; labrum sub-rectangular with lateral margins slightly convex and bend up; mandible yellowish brown with black tip, with a pair of dorsal ridges, with two large teeth ( Fig. 29 View Figs 24-32 ) and only laterally densely setose.

T h o r a x. Prothorax: pronotum evenly curved at sides and attenuate towards apex from basal third onwards, pronotum broadest 0.3 mm apically of base, without impressions; lateral margins even, carinate, completely visible in dorsal view, hind edges rounded, lobate; pronotum with puncturation regularly dense and shallow, disc with longitudinal shallow wrinkles, prothorax densely covered with fine punctures, front margin concave, lateral margins slightly projecting apically, hind margin deeply concave; mesothorax: scutellum smooth, black; elytra basally broadest then gradually attenuate, punctures small and in nine regular rows plus scutellar row and row abutting lateral margin of elytra, punctures becoming indistinct in rows eight and nine, all striae with short white adhesive setae, interstices 6-10 convex, glabrous; metathorax: legs yellowish brown, with apical 2/3 of tibia and tarsi black, claw-segment apically yellowish brown, claws black, appendiculate, fore tibiae slightly bend.

A b d o m e n Sternites yellowish brown, with short white setae, sternite VII with a shallow oval fovea, tergites and pygidium brown, sternite III as long as sternites IV to VII along midline; aedeagus ( Figs 16, 17 View Figs 14-23 ), length of the aedeagus: 1.6 mm.

D e s c r i p t i o n o f f e m a l e ( Figs 42 View Figs 33-43 , 50 View Figs 49-52 ). Dorsum brown with yellow markings as in male; venter including pygidium yellowish brown except for some blurred brownish darkenings on metathorax, and mesothorax yellow; base of pygidium with an indistinct ridge; mandibles, clypeus and labrum simple; spermatheca hook-shaped, 0.4 X 0.5 mm, spermathecal ductus straight, base simple, strongly coiled up close to spermatheca ( Fig. 42 View Figs 33-43 ); size [mm]: length 5.5, width of elytra at humeri 3.7, length of elytra 3.9; length of pronotum 2.0, maximum width 3.8.

V a r i a b i l i t y. Anteriormarginoffemalepronotum at middle dark brown or yellowish brown, variability in size [mm] (mean±SD (max., min., n)): length of female 5.82±0.17 (5.90, 5.60, 6), width of elytra at humeri 3.99±0.16 (4.15, 3.80), length of pronotum 2.05±0.05 (2.10, 2.00) and width 3.97±0.08 (4.10, 3.90).

D i f f e r e n t i a l d i a g n o s i s.Differsfromtheotherspecieswithsetoseelytra and black tarsi, Scaphodius amieus nov.sp., by the presence of lateral strongly contrasting yellow spots on the elytra, and in the male by the concave clypeus and the large, quadrate labrum.

D i s t r i b u t i o n a n d b i o l o g y. Known from the North of New Caledonia mainland only.

E t y m o l o g y. Mega means large, gnathos is Greek for jaw, referring to the male mandibles.

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Chrysomelidae

Genus

Scaphodius

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