Efesus Simutnik, 2020

Simutnik, S. A., Perkovsky, E. E. & Vasilenko, D. V., 2020, Efesus trufanovi Simutnik gen. et sp. n. (Hymenoptera: Chalcidoidea: Encyrtidae) from late Eocene Danish amber, Russian Entomological Journal 29 (3), pp. 298-302 : 299

Genus Efesus Simutnik , gen.n.

Type species. Efesus trufanovi Simutnik , sp.n.

DIAGNOSIS. Row of filum spinosum double, long, consisting of 9 setae; clava large, with rounded apex, as long as 5 preceding flagellomeres combined; notauli very short; parastigma only slightly widened; cerci advanced almost to middle of metasoma.

DESCRIPTION of female. Habitus as in Figs 1–3 . Body compact, not flattening. Head slightly wider than thorax in dorsal view and approximately rounded in frontal view, hypognathous; occipital margin rounded; surface of frontovertex with polygonal reticulate sculpture, elongated on genae, without deep punctures ( Fig. 1 ); ocelli large, in approximately right triangle; distance between posterior ocellus and eye margin (OOL) as posterior ocellar diameter; genae long, height of eye slightly larger than malar space ( Fig. 2 ); scrobal depression deep, with sharp apical margin and small interantennal prominence, distance between apical margin of scrobal depression and anterior ocellus equal to POL ( Fig. 6 ); eye bare, without hairsprings. Antenna 11-segmented (1:1:6:3); antennal scape slightly widened and flattened; pedicel conical, longer than first two flagellomeres; flagellum not flattened or widened, width of flagellomeres increases toward apex; F1and F2 small, F1–F5 almost square, F6 slightly longer than width; clava with rounded apex, 3- segmented, slightly wider than last flagellomere, about as long as 5 preceding flagellomeres combined; flagellum and clava covered with short hairs; at least F4–F6 and all 3 claval segments with mps ( Fig. 6 ). Mesosoma . Pronotum in dorsal view short, transverse; mesoscutum wider than long, with very short notauli ( Fig. 1 (n)); dorsum and ventrum with same polygonal reticulate sculpture; axillae large, with meeting inner angles; mesoscutum approximately as long as scutellum, convex; mesopleuron similarly sculptured; propodeum short. Appendages. Forewing without dark areas, hyaline; linea calva with 9 closing setae (filum spinosum) along distal margin and a row of long covering setae along proximal margin ( Fig. 4 , fs); submarginal vein with slightly widened parastigma ( Fig. 4 , p) and hyaline break; marginal vein 2.5 times as long as wide; stigmal vein straight, not curved, slightly longer than marginal vein, with distinct uncus; postmarginal vein 3 times as long as marginal vein; setae of marginal fringe short; coxae and legs with same reticulate sculpture, as thorax; tarsi five segmented; protibia with curved tibial spur; mesotibial spur thick, as long as mid basitarsus. Metasoma slightly shorter than mesosoma; cerci advanced almost to middle of metasoma ( Fig. 5 , c); 1–3 terga of gaster (Mt2–Mt4) direct and parallel, Mt5–Mt7 M-shaped, Mt8 Vshaped; all terga and sterna with polygonal reticulate sculpture; hypopygium transverse, not reaching apex of gaster ( Fig. 3 , hyp); ovipositor short, slightly protruding.

Male unknown.

SPECIES COMPOSITION. Type species.

ETYMOLOGY. “ Efes ” from fs – filum spinosum, with suffix “us”. Gender masculine.

COMPARISON AND REMARKS. The new genus differs from all known fossil genera by the double row of filum spinosum and almost not expanded parastigma. Among the extant representatives, the double and long row of closing setae also rare. For example, in Coenocercus puncticollis Thomson the filum spinosum also consists of 2 rows of setae [ Simutnik 2010, Fig. 1 ].

Figures

Figs 1–6. Efesus trufanovi, gen. et sp.n., holotype ♀: 1–3 — habitus:1, dorsal (n — notaulus); 2 — lateral (c — cercus); 3 — posterodorsal (hyp — hypopygium); 4 — forewing (fs — filum spinosum, p — parastigma, u — uncus); 5 — metasoma (Mt8 — last metasomal tergum); 6 — antennae (mps — multiporous plate sensilla), head, mesoscutum, mesotibial spur (s). Рис. 1–6. Efesus trufanovi, gen. et sp.n., голотип ♀: 1–3 — обЩий вид: 1 — сверху (n — нотаулЯ); 2 — сбоку (c — церка (пигостиль)); 3 — сЗади и сбоку (hyp — гипопигий); 4 — переднее крыло (fs — filum spinosum, p — парастигма, u — ункус); 5 — метасома (Mt8 — последний тергит); 6 — усики (mps — multiporous plate sensilla), голова, Щит среднеспинки, шпора средней голени (s).