Juraheterosmylus antiquatus, Wang, Yongjie, Liu, Zhiqi, Ren, Dong & Shih, Chungkun, 2010

Wang, Yongjie, Liu, Zhiqi, Ren, Dong & Shih, Chungkun, 2010, A new genus of Protosmylinae from the Middle Jurassic of China (Neuroptera: Osmylidae), Zootaxa 2480, pp. 45-53 : 48-49

Juraheterosmylus antiquatus sp. nov.

( Figs. 1 A–B, 2A–B)

Diagnosis. Cross-veins r1-rs and outer gradate series edged with fuscous spots.

Description. Head deformed, quadrate; eyes protruded, distinctly dark; antennae not preserved. Prothorax wider than long, quadrate; mesothorax bulging; metathorax short, with full-developed sclerites. Abdomen 10 segmented; first segment reduced, caudal segment approximately pyriform, resembling the ectoproct of extant osmylids.

Forewing 23.4mm long (right forewing and hindwing overlapped, and apex of left forewing not preserved), 7.2mm wide. Membrane with few spots, pterostigma light yellow; nygmata brown, present at the base between Rs and MP; trichosors present in the anterior margin and posterior margin; cross-veins r1-rs and outer gradate series edged with fuscous spots. Venation: costal cross-veins simple, dense close to the apex, with few distal forks; Sc and R1 fused distal; cross-veins r1-rs six branches; Rs originate from R1; Rs with 13 branches, each branch sinuous in the middle, and forming complicate distal forks; cross-veins between Rs branches arranged regularly, forming three rows of gradate series; MP forked close to the separation of MA from Rs, only four cross-veins present between both branches; Cu forked at the wing base, both branches approximately with equal length; CuA with four distal branches, CuP with three distal branches; A1 with four distal branches, its length beyond half of CuP; A2 with two simple branches; A3 simple. Hindwing 22.8mm long (incomplete, as preserved), 7.3mm wide. Membrane resembling the forewing, with few spots; pterostigma light yellow; nygmata not detected due to the poor preservation; trichosors present along the anterior and posterior margin. Venation: costal cross-veins simple, with few distal forks; r1-rs cross-veins five branches; Rs stem with a short, basal additional vein; Rs with 10 branches; Rs region poorly preserved; MP forked close to the wing base, MP with complicated distal forks; Cu forked at wing base, both branches and anal region preserved poorly.

Holotype. CNU-NEU-NN2010201. Specimen consists of a relatively complete body with wings stretched-out, but the right forewing and hindwing are overlapped.

Type locality. Daohugou Village, Shantou Township, Ningcheng County, Inner Mongolia, China.

Type horizon. Jiulongshan Formation, Bathonian–Callovian boundary, Middle Jurassic.

Etymology. The specific name is from the Latin word antiqutus (meaning, old), referring to the oldest record of Protosmylinae.

Figures

FIGURE 1. Photograph and reconstruction of new species. A – B, Juraheterosmylus antiquates gen. et sp. nov. Body with three stretched-out wings. C – D, Juraheterosmylus astictus sp. nov. Body with four stretched-out wings poorly preserved. E – F, Juraheterosmylus minor sp. nov. Complete body with wings stretched-out, but wings are poorly preserved.