Dzhungarocosa omelkoi, Marusik, Yuri M., 2017

Marusik, Yuri M., 2017, A survey of East Palaearctic Lycosidae (Araneae). 13. A new genus of spiny-legs Pardosinae from Eastern Kazakhstan, Zootaxa 4320 (2), pp. 339-350 : 345-348

Dzhungarocosa omelkoi sp. n.

Figs 5–8 , 16–18 , 25–27 , 35–38 , 49–51 , 57–61 , 69–71 , 77–80

Types. Holotype ♂ (ISEA, 001.7316) and paratypes 1♂ 3♀ (ISEA, 001.7317) KAZAKHSTAN, Almaty Region, watershed of Ermenesai and Koksai Rivers (44°43’N, 79°19’E), stony alpine meadow with screes, 2900–3000 m, 3.07.2016 (A.A. Fomichev). Paratypes: 3♂ 2♀ (ZMMU) same locality (44°43’N, 79°18’E), mountain stony tundra with screes, 3000–3100 m, 3.07.2016 (A.A. Fomichev).

Etymology. The specific name is a patronym in honour of our friend and colleague Mikhail M. Omelko (Vladivostok, Russia), for his noteworthy contribution to the study of Acantholycosa and related genera.

Diagnosis. The new species differs from the sibling D. ballarini sp. n. by smaller size and shape of copulatory organs. Males of D. omelkoi sp. n. have no digitiform anterior paleal process, present in D. ballarini sp. n. ( Figs 47, 50 ). The two species also differ by the shape of terminal apophysis, which is more massive in D. omelkoi sp. n. ( Figs 46–48, 49–51 ). Females of the two sibling species differ by the proportion of septal base, which is as long as wide in D. omelkoi sp. n. vs wider than long in D. ballarini sp. n. ( Figs 57, 62 ); by the distance between septal pockets (touching each other in D. omelkoi sp. n. vs spaced in D. ballarini sp. n., Figs 60, 65 ) and the shape of posterior part of septal base ( Figs 61, 66 ).

Description. Male. Total length 8.7. Carapace: 4.0 long, 3.15 wide. Colouration ( Fig. 5 ): carapace, sternum and labium black. Clypeus yellow with a black stripe on its lower edge. Chelicerae black with yellow stripes. Endites brown-gray with a yellow spot on the inner side. Palps: Fe, Ti and cymbium black, Pt yellow. Legs: I and II: Fe dark-gray, yellow apically, Pt–Ta yellow; III and IV yellow-gray. Abdomen black, without any pattern. Leg I joints: 3.65+1.6+3.55+3.85+1.65. Spination of leg I: Fe: d1-1-1 p0-1-3 r0-1-1; Pt: d1-1 p1-0 r1-0; Ti: d1-0-1 p1-0- 0 r1-0-0 pv6 rv6; Mt: p1-1-0 r1-1-0 v2-2 -0.

Palp as in Figs 16–18 , 25–27 , 35–38 , 49–51 . Tegular apophysis with large, claw-like anterior arm; anterior process of palea (Ap) broad, almost straight; posterior process (Pp) with 2 outgrowths on the lateral edges; tip of paleal conductor rounded; terminal apophysis massive, with rounded outgrowths; embolus smoothly bent (wavy).

Female. Total length 8.0. Carapace: 4.0 long, 3.2 wide. Colouration ( Fig. 6 ): carapace black. Clypeus yellow. Chelicerae yellow basally and brown apically. Sternum and labium dark-gray. Endites yellow-gray. Legs and palps gray with yellow rings. Abdomen dark-gray dorsally, covered by shining hairs and yellow-gray ventrally. Leg I joints: 3.6+1.7+3.3+3.1+1.45. Spination of leg I: Fe: d1-1-1 p0-1-2 r0-1-1; Pt: d1-1; Ti: d1-0-1 p1-0-0 r1-0-0 pv6 rv6; Mt: p0-1-0 r0-1-0 v2-2 -0.

Epigyne as in Figs 57–61 , 69–71 . Septum with thin stem, base of septum as wide as long; septal pockets located posteriorly, touching each other ( Fig. 60 ), base of septum in posterior view ( Fig. 61 ) longer than wide; head of receptacles converging.

Size variation. Males vary from 7.5 to 9.4 in body length, carapace 3.7–4.1 long and 3.0–3.35 wide (n=5). Females vary from 8.0 to 9.4 in body length, carapace 4.0–4.3 long and 3.2–3.55 wide (n=5).

Distribution. Type locality only ( Figs 79–80 ).

Figures

FIGURES 1 – 12. Habitus and retrolateral view of leg I in Dzhungarocosa zhishengi sp. n. (1 – 4), D. omelkoi sp. n. (5 – 8) and D. ballarini sp. n. (9 – 12). 1, 5, 9 male, dorsal; 2, 6, 10 female, dorsal; 3, 7, 11 male leg I; 4, 8, 12 female leg I. Scale = 2 mm for 1 – 2, 5 – 6, 9 – 10; 1 mm for 3 – 4, 7 – 8, 11 – 12.

FIGURES 13 – 21. Male palp of Dzhungarocosa zhishengi sp. n. (13 – 15), D. omelkoi sp. n. (16 – 18) and D. ballarini sp. n. (19 – 21). 13, 16, 19 ventral; 14, 17, 20 retrolateral; 15, 18, 21 tegulum and tegular apophysis, ventral. Scale = 0.2 mm if not otherwise indicated. Abbreviations: Aa anterior arm of tegular apophysis, Pa posterior arm of tegular apophysis, Tc tegular conductor.

FIGURES 22 – 30. Bulb and embolic division of Dzhungarocosa zhishengi sp. n. (22 – 24), D. omelkoi sp. n. (25 – 27) and D. ballarini sp. n. (28 – 30). 22, 25, 28 bulb, ventral; 23, 26, 29 bulb, retrolateral; 24, 27, 30 embolic division, ventral. Scale = 0.2 mm.

FIGURES 31 – 42. Bulb of Dzhungarocosa zhishengi sp. n. (31 – 34), D. omelkoi sp. n. (35 – 38) and D. ballarini sp. n. (39 – 42). 31, 35, 39 embolic division, apical; 32, 36, 40 embolic division, prolateral; 33, 37, 41 embolic division, retrolateral; 34, 38, 42 tegulum and tegular apophysis, apical. Scale = 0.2 mm.

FIGURES 43 – 51. Embolic division of Dzhungarocosa zhishengi sp. n. (43 – 45), D. ballarini sp. n. (46 – 48) and D. omelkoi sp. n. (49 – 51). 43, 46, 49 embolic division, ventral; 44, 47, 50 embolic division, apical; 45, 48, 51 embolic division, ventroretrolateral. Scale = 0.1 mm. Abbreviations: Ap anterior process of palea, Co conductor, Em embolus, Pp posterior process of palea, Rp rounded pit of embolic division, Ta terminal apophysis.

FIGURES 52 – 66. Epigyne of Dzhungarocosa zhishengi sp. n. (52 – 56), D. omelkoi sp. n. (57 – 61) and D. ballarini sp. n. (62 – 66). 52, 57, 62 ventral; 53, 58, 63 macerated, ventral; 54, 59, 64 macerated, dorsal; 55, 60, 65 anterior; 56, 61, 66 posterior. Scale = 0.2 mm. Abbreviations: Li lip of the epigyne, Rh head of the receptacle, Po septal pocket.

FIGURES 67 – 74. Epigyne of Dzhungarocosa zhishengi sp. n. (67 – 68), D. omelkoi sp. n. (69 – 71) and D. ballarini sp. n. (72 – 74). 67, 69, 72 ventral; 68, 70, 73 ventro-lateral, showing septal pocket; 71, 74 posterior. Scale = 0.1 mm (if indicated). Abbreviations: Bs base of septum, Ea sclerotized arches, Po posterior pockets, St septal stem.

FIGURES 75 – 80. Habitats and type localities of Dzhungarocosa zhishengi sp. n. (75, 79, 80 — square), D. ballarini sp. n. (76, 79, 80 — circle) and D. omelkoi sp. n. (77 – 79, 80 — star).