Amplaria rykkenae, Shear, 2021

Shear, William A., 2021, The millipede family Striariidae Bollman, 1893. II. New records and species of the genus Amplaria Chamberlin, 1941 (Diplopoda, Chordeumatida, Striarioidea), Zootaxa 4908 (2), pp. 205-224 : 212

Amplaria rykkenae , new species

Figs. 11–14 , 51, 52

Types: Male holotype from 1 mi ENE of Upper Soda, Sweet Home Road, Willamette National Forest , 44.4131°, -122.2619°, 1840’ asl, Linn Co. Oregon, collected 1 October 2001 by Jessica Rykken; male paratypes from 4 mi WSW of Upper Soda, 44.4015°, -122.3749°, 1440’ asl, collected 27 November 2001, and from 4 mi WNW of Upper Soda, 44.4085°, -122.3844°, 612’ asl, collected 15 August 2001, both localities on Sweet Home Road, Willamette National Forest , Linn Co. , Oregon, both collected by Jessica Rykken. Parts of these specimens are mounted on SEM stub WAS34-7 .

Etymology: I am pleased to name this species for the collector, Jessica Rykken, presently entomologist at Denali National Park, Alaska.

Diagnosis: Distinct from A. arcata , the other species in the group, in having the femora of the second legpair in males of normal size, and the more prominent acute process at the tip of the gonopod anterior angiocoxite ( Fig. 12 ).

Description: Holotype male. Length, about 11 mm, width about 1.0 mm. Seven or 8 poorly formed, unpigmented ommatidia in triangular patch. Labral hooks long, nearly straight. Mandible not modified. Legpairs 1 and 2 similar in size, first legs without long setae, tarsi ventrally armed with twisted, spatulate setae. Trochanters of second legs without processes, small group of specialized setae present, femora of normal size. Coxal flasks of third legs evenly curved, tapering, pleurotergal bars of third ring narrow, not meeting in midline. Legpairs 4–7 enlarged, with flattened podomeres, coxae unmodified except for lobes on coxae 7. Metazonital crests moderate, segmental setae long, filamentous. Pygidium elongate, deeply divided between lobes ( Fig. 14 ). Color uniform pale tan.

Gonopods ( Figs. 11, 12 , 51, 52 ) with numerous spiculate processes posterioapically on anterior angiocoxites, coxites cup-like apically; posterior angiocoxites rather narrow, not greatly expanded distally, sheathing 3 or 4 flagellocoxites. Ninth legpair ( Fig. 13 ) with long lateral coxal apophysis, mesally with poorly sclerotized fungiform area possibly representing vestigial coxal glands.

Females not collected.

Distribution: Known only from the general vicinity of Upper Soda, Oregon.

Figures

FIGS. 11–13. Amplaria rykkenae. Fig. 11. Gonopods, anterior view. Fig. 12. Gonopod tips, anterior view. Fig. 13. Legpair 9, anterior view.

FIGS. 14–19. Amplaria rykkenae, A. arcata. Fig. 14. Amplaria rykkenae, posterior end of male, dorsal view. Figs. 15–19. A. arcata. Fig. 15. Left side of head and first two legpairs of male, anterior view. Fig. 16. Tarsus of legpair 1, lateral view. Fig. 17. Special setae of tarsus 1. Fig. 18. Second and third legpair of male, ventral view. Fig. 19. Pygidium, ventral view.

FIGS. 45–54. Drawings of gonopods of Amplaria species. Fig. 45. Left gonopod of A. nazinta, lateral view. Fig. 46. Tip of anterior angiocoxite of A. nazinta gonopod, lateral view. Fig. 47. Right gonopod of A. crawfordi, mesal view. Fig. 48. Tip of anterior angiocoxite of A. crawfordi, lateral view. Fig. 49. Right gonopod of A. fontinalis, mesal view. FIG. 50. Tip of anterior angiocoxite of A. fontalis, posteriolateral view. Fig. 51. Right gonopod of A. rykkenae, mesal view. Fig. 52. Right gonopod of A. rykkenae, lateral view. Fig. 53. Right gonopod of A. baughi, lateral view. Fig. 54. Right gonopod of A. umatilla, lateral view.