Amplaria crawfordi, Shear, 2021

Shear, William A., 2021, The millipede family Striariidae Bollman, 1893. II. New records and species of the genus Amplaria Chamberlin, 1941 (Diplopoda, Chordeumatida, Striarioidea), Zootaxa 4908 (2), pp. 205-224 : 209

Amplaria crawfordi , new species

Figs. 4–6 , 47, 48

Types: Male holotype and male and female paratypes from west of Thornton Creek, 47.698°, -122.279°, 80’ asl, Seattle , King Co., Washington, collected 10–15 April 2002 by Jerry Austin ; deposited in Burke Memorial Museum of the University of Washington, Seattle , Washington. Parts of these specimens are mounted on SEM stub WAS34-8 .

Diagnosis: Similar to and clearly related to A. nazinta and A. fontinalis , n. sp. Distinct from the former in the longer, finger-like extension of the ninth leg telopodite ( Fig. 4 ), and from the latter in having a few more ommatidia and in details of the gonopod (compare Figs. 2 and 5 ).

Etymology: I take pleasure in naming this species for my esteemed colleague, Rod Crawford, curator of terrestrial invertebrates at the Burke Memorial Museum of the University of Washington, Seattle, Washington.

Description: Male holotype. Length, about 10 mm, width 0.9–1.0 mm. Four small, poorly formed black ommatidia. Labrum without hooks; mandible not modified. First legpair larger than second and third, femur and postfemur with long, needle-like setae. Second legpair coxae fused in midline and to their respective trochanters, short apophysis with specialized setae on trochanteral part ( Fig. 6 ). Third coxal flasks basally fused, distally tightly appressed in midline, enlarged, sharply angled anteriorly; third telopodites reduced in size, prefemur with strong, distoventral lobe ( Fig. 6 ). Pleurotergal bars of third ring broad, not touching in midline. Coxae of legpairs four and five not modified, telopodites enlarged, with flattened podomeres. Pygidium short. Color medium chestnut brown (after long preservation).

Gonopods ( Fig. 5 , 47, 48 ) with anterior angiocoxite apically divided; lateral branch slender, rebranched, deflexed at tip. Posterior angiocoxite expanded, in situ sheathing three or four flagellocoxites. Ninth legpair coxae without prominent apophysis, mesal apex acute; telopodite with long, finger-like mesal extension ( Fig. 4 ).

Females similar in non-sexual characters to males.

Distribution: Known only from the type locality.

Figures

FIGS. 1–4. Amplaria nazinta, A. crawfordi. Figs. 1–3. Amplaria nazinta. Fig. 1. Rings 8 and 9, lateral view. Fig. 2. Gonopods, anterior view. Fig. 3. Legpair 9, anterior view. Fig. 4. A. crawfordi, right leg 9, anterior view.

FIGS. 5–7. Amplaria crawfordi, A. fontinalis. Figs. 5, 6. Amplaria crawfordi. Fig. 5. Gonopods, anterior view. Fig. 6. Legpairs 2 and 3, ventral view. Fig. 7. A. fontinalis, left gonopod, lateral view.

FIGS. 45–54. Drawings of gonopods of Amplaria species. Fig. 45. Left gonopod of A. nazinta, lateral view. Fig. 46. Tip of anterior angiocoxite of A. nazinta gonopod, lateral view. Fig. 47. Right gonopod of A. crawfordi, mesal view. Fig. 48. Tip of anterior angiocoxite of A. crawfordi, lateral view. Fig. 49. Right gonopod of A. fontinalis, mesal view. FIG. 50. Tip of anterior angiocoxite of A. fontalis, posteriolateral view. Fig. 51. Right gonopod of A. rykkenae, mesal view. Fig. 52. Right gonopod of A. rykkenae, lateral view. Fig. 53. Right gonopod of A. baughi, lateral view. Fig. 54. Right gonopod of A. umatilla, lateral view.