Amplaria staceyi, Shear, 2021

Shear, William A., 2021, The millipede family Striariidae Bollman, 1893. II. New records and species of the genus Amplaria Chamberlin, 1941 (Diplopoda, Chordeumatida, Striarioidea), Zootaxa 4908 (2), pp. 205-224 : 215-216

Amplaria staceyi , new species

Figs. 31–34 , 58, 59

Types: Male holotype and male and female paratypes from the Weir Trail, Palomar Mountain State Park, 33.3435°, -116.9072°, 1395 m asl, San Diego Co., California, collected March 26, 2017, by C. Richart, BJ Stacy, C. Lee, M. Salkiewicz & C. DeGroof (Richart collection #CHR5309). Additional paratypes from Pauma Valley , Palomar Mountain , 33.3417°, -116.9044°, collected 12 February 2017 by C. Richart, BJ Stacey & J. Keller (Richart collection #CHR5216), and from South Grade Road , Palomar Mountain , 33.3065°, -116.8774°, 13 22 m asl, collected 5 May 2011 by A. Schoenhofer. Parts of these specimens are mounted on SEM stub WAS34-12 .

Diagnosis: Very close to A. imberbis , but distinct in details of the gonopods (cf. Figs. 33–34 , 58, 59 with Figs. 11, 12 in Shear [2020]).

Etymology: As requested by Casey Richart, this species is named for BJ Stacey for his extraordinary contributions to community science and for being among the collectors that discovered this species.

Description: Male holotype. Length about 9.0 mm, width 0.85 mm. Five black ommatidia in triangular patch. Labrum without hooks. Mandibular stipes with acute distal corner, otherwise not modified. Legpair 1 larger than legpair 2, without needle-like setae, tarsus with twisted, spatulate setae ventrally. Trochanters of legpair 2 with ventral projection about twice as long as podomere, densely set with specialized setae ( Fig. 32 ). Legpair 3 coxal flasks short, about as long as width of coxae; pleurotergal bars of ring 7 short, scarcely reaching lateral sides of third coxae ( Fig. 31 ). Legpairs 4–7 enlarged, podomeres flattened. Metazonital crests of moderate height. Pygidium short. Color pale tan with “2-stripe” pattern in darker purplish brown, purplish brown pleural spots, pygidium white.

Gonopods ( Figs. 33, 34 , 58, 59 ) simple, anterior angiocoxite scoop-like, with small, triangular lateral tooth subapically; posterior angiocoxite basally narrow, sharply curved, apically widened, sheathing four acute flagellocoxites. Ninth legpair without coxal process, telopodite typical.

Females similar to males in nonsexual characters.

Distribution: Known from several collections from Palomar Mountain.

Figures

FIGS. 29–31. Amplaria baughi, A. staceyi. Figs. 29, 30. Amplaria baughi. Fig. 29. Left leg 3 of male, anterior view. Fig. 30. Right leg 9 of male, anterior view. Fig. 31. A. staceyi, lepair 3 of male, posterioventral view.

FIGS. 32–36. Amplaria staceyi, A. umatilla. Figs. 32–34. Amplaria staceyi. Fig. 32. Apophysis of left leg 2 of male, ventral view. Fig. 33. Left gonopod of male, lateral view. Fig. 34. Right gonopod of male, mesal view. Figs. 35, 36. A. umatilla. Fig. 35. Second and third legpair of male, ventral view. Fig. 36. Gonopod complex, lateral view.

FIGS. 55–59. Drawings of gonopods of Amplaria species. Fig. 55. Right gonopod of A. umatilla, mesal view. Fig. 56. Right gonopod of A. cervus, mesal view. Fig. 57. Right gonopod of A. mendocino, mesal view. Fig. 58. Right gonopod of A. flucticulus, lateral view. Fig. 59. Right gonopod of A. flucticulus, mesal view.

FIGS. 11–13. Amplaria rykkenae. Fig. 11. Gonopods, anterior view. Fig. 12. Gonopod tips, anterior view. Fig. 13. Legpair 9, anterior view.