Tonnoiriella syriensis Ježek, 1999

Wagner, Rüdiger & Withers, Phil, 2020, The West-Palearctic species of the genus Tonnoiriella Vaillant, 1971 (Diptera: Psychodidae, Psychodinae), Zootaxa 4728 (2), pp. 183-210 : 196-197

Tonnoiriella syriensis Ježek, 1999

( Figs. 24, 25 )

Tonnoiriella syriensis Ježek, 1999 : Časopis Národniho Muzea v Praze Řada přirodovedna 168: 14.

Material: 1♂ holotype, slide mounted: head with mouthparts 2 palpi, 1 antenna, thorax cut in half, 2 wings, 4 legs, remains of abdomen, styli with aedeagus, epandrium with epandrial processes; left side white label: Syria 8303, Al Lathiqiyeh Province , NW of Rabi’ah (35°49’N / 36°02’E) 2.7.1998, a stream approximately 550 m a.s.l. Chvoika leg., near a forest brook with close undergrowth. Right side red label: Tonnoirella syriensis Ježek 1999 , holotype Cat. No. 33845 male 1999. Paratypes: Cat. Nos. 33846, and 33847, both collected with holotype (in Natural History Museum Prague) .

1♂ (GBIFCH 00585194), Cyprus, 6 km E of Panagia , 34°55’N / 32°42’E, 550 m a.s.l; small stream, water temperature 16.4°C, 2 May 1974 ; 1♂, (GBIFCH 00585192), Cyprus, 7 km E of Panagia , 34°55’N / 32°42’E, 600 m a.s.l; small stream, water temperature 16.0°C, 2 May 1974 ; 1♂, (GBIFCH 00585190), Cyprus, Paliambela , 35°08’N / 32°35’E, 130 m a.s.l; left stream, above dam, 17.9°C, 5 May 1974 , all leg. Malicky (all in the collection F. Vaillant deposited in the Natural History Museum Lausanne).

Description: head with eyebridge of 3 facet rows; distance between eyes 3 facet diameters; postocular bristles present. Antenna (from paratype no 33846) with scape, pedicel and 14 flagellomeres: absolute length 0.044 -0.044 - 0.03-0.032-0.034-0.034-0.036-0.036-0.032-0.032-0.028-0.03 -0.028 -0.022 - 0.02-0.028 mm; relative length 22-22- 15-16-17-17-18-18-16-16- 14-15-14-11 -10-14; terminal flagellomere with apiculus. Pairs of digitate ascoids on flagellomeres 5–11. Palpus segments absolute length: 0.048-0.050-0.066-0.142; relative length 24-25-33-71.

Wing 2.85 times longer than wide; length 1.46 mm, width 0.51 mm.

Male terminalia ( Fig. 24 ) with median part of hypandrium not clearly discernable, probably equally thin. Gonocoxites 2x longer than wide, with a long seta at about middle; basal group of setae on inner side loosely grouped. Gonostyli 1.3 times longer than gonocoxites, slightly bent. Parameres difficult to identify, small.

Aedeagus with a comparatively short basiphallus; ventral distiphallus with ventral arm apically rounded Tshaped, dorsal arm less than half as long, straight. Dorsal lobe and distiphallus are alike T-shaped straight, the cross sclerite shorter than its ventral couterpart.

Epandrium rectangular; epandrial processes almost straight with 4 tenacula, apical two about 1/3 the length of the basal ones ( Fig. 25 ).

Specimens from Cyprus differ from typical specimens by the distance between eyes (4 facet diameters), wing features (3.04 times longer than wide; length 1.84 mm, width 0.61 mm).

Distribution: Syria, Cyprus.

Remarks: Species well distinguishable from congeners due to the shape of the distiphallus bilobed on both sides. The shape of the median part of the hypandrium is not clearly discernable. The genitalia of the holotype from Syria and the males from Cyprus are identical. Minor differences occur in wing size and length of palpus and antenna segments, which are in the size variation of species.

Figures

FIGURES 15–25: 15 T. sieberti genitalia ventral view; 16 T. ikariae genitalia ventral view; 17–21 T. italiae, genitalia ventral view of specimen from northern Italy (17), Sicily (18), aedeagus of specimen from N Italy (19), Sicily, Villa doso (20) and Sicily Castelbuono (21); 22 T. turcica genitalia ventral view; 23 T. rhodesica genitalia ventral view; 24 T syriensis genitalia ventral view, 25 T. syriensis epandrium and epandrial processes oblique view.