Tonnoiriella sieberti Wagner, 1993

Wagner, Rüdiger & Withers, Phil, 2020, The West-Palearctic species of the genus Tonnoiriella Vaillant, 1971 (Diptera: Psychodidae, Psychodinae), Zootaxa 4728 (2), pp. 183-210 : 192

Tonnoiriella sieberti Wagner, 1993

( Fig. 15 )

Tonnoiriella sieberti Wagner, 1993 : Entomofauna 14 (24): 407.

Material: 1♂ holotype, North Macedonia, Skopje, April 1976 ; 1♂ paratype, collected with holotype, leg. Siebert ( RW) .

Description: Head round, eyebridge of 3 facet rows; distance between the eyes 5 facet diameters; 6 or 7 postocular bristles. Antenna with scape, pedicel and 14 flagellomeres; absolute length 0.044-0.056-0.044-0.048- 0.054-0.048-0.05-0.05-0.046-0.04 -0.038 -0.042 -0.03-0.024- 0.02-0.028 mm; relative length 22-28-22-24-27-24-25- 25-23-20- 19-21-15-12 -10-14. Flagellomeres 5–11 with a pair of digitiform curved ascoids in the distal third. Palpus segments: absolute length 0.06-0.088- 0.098-0.176 mm; relative length 30-44-49-88.

Wing translucent, 2.86 times longer than wide; length 2.18 mm, width 0.76 mm.

Male terminalia ( Fig. 15 ) with hypandrium broadened in the middle with edges sinuously curved. Gonocoxites approximately 2x longer than wide, straight; inner edge basally with a group of 5–7 short setae at about middle with a single long seta; gonostylus curved, as long as gonocoxite.

Parameres with a broad ventral bridge, lobes do not reach middle of gonocoxite; ventrally in the midst develops the aedeagus sheath with a racket-shaped brownish mark; above a pair of broad bilobed processes that do not meet in the middle; further dorsal large rhomboid condyles that form the median keel.

Basiphallus bilobed, ventral lobe bifid, ventral branch connected to the distiphallus, longer part slightly bent with blunt tip; distiphallus with both ends rounded, the broader end with a basal triangular projection, the smaller end rounded bifid. The dorsal lobe of the basiphallus is distally fixed to the anchor-shaped distiphallus with cross end, one side broad the other side a thin band, no apical foldings.

Epandrium rhomboid, epandrial processes almost straight, apically with 10–12 feathered tenacula.

Distribution: Albania, Czech Republic, France, Greece, North Macedonia, Syria.

Remarks: T. sieberti is distinguished from the other group members by the shape of the median hypandrium, the large ventral lobes of the parameres, and the apically rounded ventral distiphallus. Ježek (1999) mentioned the species from the Czech Republic, North Macedonia, Albania, Greece and Syria. However, it is possible that in Greece and in particular in Syria other species occur.

Figures

FIGURES 15–25: 15 T. sieberti genitalia ventral view; 16 T. ikariae genitalia ventral view; 17–21 T. italiae, genitalia ventral view of specimen from northern Italy (17), Sicily (18), aedeagus of specimen from N Italy (19), Sicily, Villa doso (20) and Sicily Castelbuono (21); 22 T. turcica genitalia ventral view; 23 T. rhodesica genitalia ventral view; 24 T syriensis genitalia ventral view, 25 T. syriensis epandrium and epandrial processes oblique view.