Tonnoiriella ikariae, Wagner & Withers, 2020

Wagner, Rüdiger & Withers, Phil, 2020, The West-Palearctic species of the genus Tonnoiriella Vaillant, 1971 (Diptera: Psychodidae, Psychodinae), Zootaxa 4728 (2), pp. 183-210 : 194

Tonnoiriella ikariae sp. nov.

( Fig. 16 )

Material: 1♂, holotype, Greece, Ikaria island , 2 km East of Karavostomou (37°38’N / 26°13’E), spring, 180 m a.s.l, 30 May 1979, leg. Malicky ( RW) .

Description: Head with an eyebridge of 3 facet rows; distance between eyes 4 facet diameters; several postocular bristles behind eyes. Antenna with scape, pedicel and 14 flagellomeres: absolute length 0.038-0.05-0.042-0.044- 0.042-0.042-0.044-0.044-0.046-0.046-0.036-0.036-0.03-0.022- 0.02-0.044 mm; relative length 19-25-21-22-21-21- 22-22-23-23- 18-18-15-11 -10-12. Flagellomeres 5–11 with a pair of digitiform curved ascoids. Palpus segments, absolute length: 0.05-0.062- 0.074-0.138 mm; relative lenght 25-31-37-69.

Wing 2.70 times longer than wide; length 1.82 mm, width 0.67 mm.

Male terminalia ( Fig. 16 ) with hypandrium expanded along 1/3 of its ventral length with 2 lateral and 1 median widenings. Gonocoxite approximately 2x longer than its median width; inner side with a basal group of setae, just before middle 2 long setae; gonostylus longer than gonocoxite, almost straight.

Parameres with a broad ventral bridge, lobes just reach the bases of the long setae on gonocoxite, ventrally in the midst with a racket-shaped brownish spot of aedeagus sheath; above a pair of processes as long as the more ventral processes that do not meet in the middle. Further dorsal large basally round condyles with parallel inner edges form the keel.

Basiphallus longer than wide, basally broader, apically bilobed; ventral lobe bifid, ventral part connected to the distiphallus, longer part slightly bent with blunt tip; distiphallus longer than gonocoxites angular, with both ends acute, the broader end with a broad projection, the smaller end double-headed. The end of the dorsal distiphallus is across anchor-shaped sclerite clearly thinner than the its ventral counterpart.

Epandrium rhomboid, epandrial processes almost straight, apically with 8 feathered tenacula.

Distribution: Greece, island of Ikaria.

Etymology: The epithet is derived from the island of Ikaria, where the species was collected.

Remarks: Another species of the T. sieberti -group. The middle part of the hypandrium is less prominent than in T. turcica and T. sieberti . The shape of the hypandrium, and the long and angular distiphallus distinguish the new species from its congeners.

Figures

FIGURES 15–25: 15 T. sieberti genitalia ventral view; 16 T. ikariae genitalia ventral view; 17–21 T. italiae, genitalia ventral view of specimen from northern Italy (17), Sicily (18), aedeagus of specimen from N Italy (19), Sicily, Villa doso (20) and Sicily Castelbuono (21); 22 T. turcica genitalia ventral view; 23 T. rhodesica genitalia ventral view; 24 T syriensis genitalia ventral view, 25 T. syriensis epandrium and epandrial processes oblique view.