Tonnoiriella Vaillant 1982

Wagner, Rüdiger & Withers, Phil, 2020, The West-Palearctic species of the genus Tonnoiriella Vaillant, 1971 (Diptera: Psychodidae, Psychodinae), Zootaxa 4728 (2), pp. 183-210 : 208

Key to males of the west-palearctic species of genus Tonnoiriella Vaillant 1982

1 - Hypandrium thin without broadening, gonostyli 1.5 x longer than gonocoxites with angled tip, aedeagus with parallel sides, longer than gonocoxites, inner seta on gonocoxites shorter than the diameter of the gonocoxite...... T. graeca 1993 Wagner

- Hypandrium usually with median broadening, gonostyli about as long as gonocoxites with sharp tip, gonocoxites with a strong seta long distinctly longer than the gonocoxite diameter....................................................... 2

2 - Dorsal lobus of the basiphallus linked with a clearly detectable distiphallus sclerite................................. 3

- Dorsal lobus of the basiphallus without detectable distiphallus sclerite.......................................... 12

3 - Dorsal distiphallus extensive, lateral protrusions with sharp tips................................ T. pulchra -group ... 4

- Dorsal distiphallus small anchor-shaped, not protruding the edge of the aedeagus................... T. sieberti -group … 7

4 - Hypandrium medially with a thin tongue-shaped broadening................................. T. anderssoni (Nielsen)

- Hypandrium medially with large broadening................................................................ 5

5 - Median broadening of hypandrium concave..................................................... T. paveli Ježek

- Median broadening of hypandrium convex................................................................. 6

6 - Median broadening of hypandrium rhomboid................................................. T. pulchra (Eaton)

- Median broadening of hypandrium triangular................................................ T. aurasica sp. nov.

7 - Hypandrium thin, of equal width over the entire length.......................................... T. syriensis Ježek

- Hypandrium distinctly broadened in the midst.............................................................. 8

8 - Hypandrium broadened in the midst and less pronounced laterally............................................... 9

- Hypandrium broad over the entire ventral length, with distinct central broadening................................. 11

9 - Hypandrium broad only in the middle........................................................ T. sieberti Wagner

- Hypandrium with in addition 2 smaller lateral prominences................................................... 10

10 - Ventral distiphallus sclerite as long as gonocoxite, apically with 2 small tips.......................... T. ikariae sp. nov.

- Ventral distiphallus sclerite shorter than gonocoxite, apically blunt................................. T. turcica Wagner

11 - Hypandrium ventrally only slightly broadened................................................. T. egeica sp. nov.

- Hypandrium ventrally in the midst shield-like broadened......................................... T. italiae sp. nov.

12 - Dorsal lobus of basiphallus apically dualistic branched....................................................... 13

- Dorsal lobus of basiphallus not branched.................................................................. 14

13 - Median broadening of hypandrium 2x longer than broad...................................... T. atlantica (Satchell)

- Median broadening of hypandrium inconspicuous................................. T. fontinalis Wagner & Salamanna

14 - Gonostylus apically apruptly thinner, digitate or harpoon shaped with serrate outer edge............. T. mollis -group … 15

- Gonostylus evenly bent, tapered, epandrial processes appendages elongate with more than 5 retinacula................................................................................................. T. nigricauda -group … 17

15 - Hypandrium medially hardly broadened, epandrial processes appendages with 1 large and 2 short tenacula.................................................................................................... T. mollis (Satchell)

- Hypandrium medially linguiform broadened............................................................... 16

16 - Gonostylus as long as gonocoxite, basally globular, apically harpoon-shaped............... T. anchoriformis (Salamanna)

- Gonostylus longer than the gonocoxite, apically digitate with a line of 4 setae bases........ T. filistylis Vaillant & Moubayed

17 - Hypandrium with 2 median or lateral broadenings.......................................................... 18

- Hypandrium with 1 median broadening................................................................... 19

18 - Broadenings lateral, short, acute, ventral distiphallus sclerite almost semicircular apically acute........... T. arcuata Ježek

- Broadenings close to the middle, rounded, ventral distiphallus sclerite apically round............... T. goncalvesi sp. nov.

19 - Median broadening small, ventral distiphallus sclerite with blunt 3 lobes....................... T. nigricauda (Tonnoir)

- Median broadening distinct, ventral distiphallus sclerite with 2 sharp lobes......................... T. andradei sp. nov.