Tonnoiriella anchoriformis Salamanna, 1975

Wagner, Rüdiger & Withers, Phil, 2020, The West-Palearctic species of the genus Tonnoiriella Vaillant, 1971 (Diptera: Psychodidae, Psychodinae), Zootaxa 4728 (2), pp. 183-210 : 201

Tonnoiriella anchoriformis Salamanna, 1975

( Figs. 38–40 )

Tonnoiriella anchoriformis Salamanna, 1975 : Entomologia, Bari, 11: 208.

Tonnoiriella disneyi Withers, 1997 : Dipterists Digest 1997 (4): 61; Ježek 1999: Časopis Národniho muzea Řada přirodovědná, 168: 17 (syn.).

Tonnoiriella holmi Wagner, 1993 : Entomofauna 14 (24): 406, new synonymy.

Material: 1♂ Germany: Schleswig-Holstein, upper Schierenseebach , 17 May 1985 ; 1♂ Germany: Schleswig-Hol- stein, lower Schierenseebach , 22 May 1985 , all leg. Holm (holotype and paratype of Tonnoiriella holmi )

Description: Head round, eyebridge of 3 facet rows; distance between eyes 4.7 facet diameters; with postocular bristles. Palpus segments, absolute length: 0.072-0.076-0.080-0.152; relative length: 36-38-40-76. Antenna with scape, pedicel and 5 flagellomeres, distal segments missing: absolute length 0.044-0.052-0.040-0.040-0.038-0.036- 0.040- mm; relative length 22-26-20-20-19-18-20-; distal flagellomeres missing; flagellomere 5 and following with a pair of digitiform curved ascoids in the apical third.

Wing 2.83 times longer than wide; length 2.05 mm, width 0.72 mm.

Male terminalia with hypandrium with broad median linguiform enlargement, tip almost reaching tip of gonocoxites ( Fig. 38 ). Gonocoxites as long as their basal width, with 1 long seta near apical end on inner edge. Gonostylus with basal swelling as long as the gonocoxite incised near tip with spearhead appearance ( Fig. 39 ).

Parameres not clearly identified, small but seemingly not different from other species. Aedeagus with ventral lobe of the basiphallus short straight; distiphallus sclerite with long basal arm, cross part slightly bent, thin. Dorsal lobe of basiphallus is broad, straight, and symmetric, no dorsal distiphallus.

Epandrium rhomboid short; epandrial processes basally bulky distally thinner with 3 feathered retinacula, 1 long and 2 short ( Fig. 40 ). Hypoproct oval, microsetose.

Distribution: Italy, Germany, Great Britain.

Remarks: This well defined species is widely distributed in Western Europe. However, it is rarely collected which finally caused the repeated descriptions from Germany and England. German specimens were collected along a slow running lowland stream.

Figures

FIGURES 32–42: 32–34 T. mollis (holotype), 32 styles and hypandrium, 33 epandrial processes, lateral view, 34 epandrium, epandrial processes and proctiger; 35–37 T. mollis (paratype), 35 hypandrium, 36 epandrial processes, lateral view, 37 aedeagus dorsal view; 38–40 T. anchoriformis 38 hypandrium, gonopods, aedeagus ventral view, 39 gonostylus, 40 epandrial processes with tenacula; 41–42 T. filistylis 41 apical part of gonostylus, 42 genitalia ventral view, (figs 41 and 42 from Vaillant & Moubayed 1987).