Quercus brandegeei Goldman, Contr. U.S. Natl. Herb.

Bartholomew, Bruce & Almeda, Frank, 2023, Nomenclator botanicus of Fagaceae in Latin America, Phytotaxa 580 (1), pp. 1921-1935 : 1921-1935

Quercus brandegeei Goldman, Contr. U.S. Natl. Herb. 16: 321 (1916). Type:— MEXICO, Baja California Sur :

E. W. Nelson & E. A. Goldman 7475 (Holotype: US00089438!; isotypes: A!, NY!, UCSB (fragment)!). Distribution : Mexico (Baja California, Baja California Sur) .

Quercus breedloveana Nixon & Barrie, Novon 25(4): 444, fig. 1 (2017). Type:— MEXICO, Guerrero: J. L. Contreras 1207 (Holotype: CAS (accn. 752961)!; isotypes: CAS!, FCME, MEXU!). This species is similar to Q. cortesii , and further study is necessary to determine if they are really different.

Distribution: Mexico (Chiapas, Guerrero).

Quercus bumelioides Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 188 (1854) . Type :— PANAMA, Veraguas: B. Seemann s. n. (Feb 1849) (Holotype: K000575038 !; isotype: BM!) .

Quercus copeyensis C.H.Mull., U.S.D.A. Bur. Pl. Industr. Misc. Publ. 477: 30, pl. 31, 32 (1942). Nom. nov. for Q. costaricensis f. kuntzei Trel. (1924) non Q. kunzei Gard. (1890) .

Quercus costaricensis Liebm. f. kuntzei Trel., Mem. Natl. Acad. Sci. 20: 146, pl. 283b (1924). Type:— COSTA RICA, Cartago: O. Kuntze 2282 (Holotype: NY; isotype: K!). The specimen at US of Kuntze 2282 does not match the specimen at K or the images in Trelease (1924), pl. 283b of the NY specimen so we do not recognize it as an isotype.

Quercus pacayana C.H.Mull., U.S.D.A. Bur. Pl. Industr. Misc. Publ. 477: 30, pl. 30 (1942). Type :— GUATEMALA, Guatemala: P. C. Standley 58467 (Holotype: F0060040 F!; isotypes: F!, NA, UCSB (fragment)!).

Distribution: Costa Rica, El Salvador, Guatemala, Honduras, Mexico (Chiapas), Nicaragua, Panama.

Quercus calophylla Schltdl. & Cham., Linnaea 5(1): 79 (1830). Type:— MEXICO, Veracruz: C. J. W. Schiede 24 (Holotype at B destroyed. Lectotype (designated here): HAL0060349 !; isolectotypes: G (as s.n.)!, M!). At the time of publication of Q. calophylla, D.F.L. Schlechtendal was at B and was not transferred to HAL until 1833 (Stafleu & Cowan 1985, p.190). For this reason, the Schiede 24 holotype would have almost certainly been at B and destroyed; the specimen at HAL should be considered an isotype. It is for this reason that we are lectotypifying the specimen as HAL rather than considering it to be the holotype.

Quercus acuminata M.Martens & Galeotti non Roxb. (1832), Bull. Acad. Roy. Sci. Bruxelles 10(1): 217 (1843). See Q. calophylla f. acuminata below.

Quercus alamo Benth., Pl. Hartw. 55 (1840). Type:— MEXICO, Puebla: T. Hartweg 423 (Holotype: K000512985 !; isotypes: A!, BM!, C (as s.n.)!, CAS!, CGE!, E!, FI!, G!, L!, LD (image)!, LE!, NY!, OXF!, P!, UCSB (fragment)!).

Quercus calophylla f. acuminata Seemen ex Trel., Mem. Natl. Acad. Sci. 20: 203 (1924). Type:— MEXICO, Veracruz: H. Galeotti 96 (Lectotype: BR0000006912253! designated by Trelease (1924, p. 230, pl. 413); isolectotypes: BR!, G!, K!, P!). Quercus acuminata M.Martens & Galeotti (1843) non Roxb. (1832) is a later homonym as well as having two syntypes, but the specific epithet of M.Martens & Galeotti was used by Trelease, based on the same type which he effectively lectotypified, as the basis for Q. calophylla f. acuminata .

Quercus calophylla f. alamo (Benth.) Trel., Mem. Natl. Acad. Sci. 20: 204 (1924).

Quercus calophylla f. flavida (Liebm.) Trel., Mem. Natl. Acad. Sci. 20: 203 (1924).

Quercus calophylla f. intermedia Seemen ex Trel., Mem. Natl. Acad. Sci. 20: 204, pl. 415 (1924). Type:— MEXICO, Veracruz: H. Galeotti 132 (Holotype: BR0000006913434!; isotypes: BR!, G!, K!, P!, UCSB (fragment)!). This form is based on the later homonym Q. intermedia M. Martens & Galeotti (1843) non Boenn. ex Rchb. (1831), but is a legitimate form based on the same type.

Quercus calophylla f. schiedeana Trel., Mem. Natl. Acad. Sci. 20: 203 (1924). Type:— MEXICO, Veracruz: C. A. Purpus s. n. (Dec 1914) (Lectotype: ILL00007467 ! designated by Valencia-A. et al. (1916, p. 256)).

Quercus calophylla f. willdenovii Trel., Mem.Natl.Acad.Sci. 20: 204, pl. 411(1)(1924). Type:— MEXICO, Guanajuato: A. Bonpland 4270 (Holotype: B-W17613010!; isotypes: P!, UCSB (fragment)!).

Quercus candicans auct. non Neé f. alligata Trel., Repert. Spec. Nov. Regni Veg. 33: 319 (1934). Type:— MEXICO, Guerrero: C. Toledo & R. Landa 910 (Syntypes at B destroyed. Neotype: FCME designated by Valencia-A. et al. (2018, p. 256); isoneotype: MEXU (image)!) .

Quercus candicans auct. non Neé f. incurva Trel., Mem. Natl.Acad. Sci. 20: 205, pl. 418(2) (1924). Type:— MEXICO, Jalisco: J. N. Rose 2999 (Lectotype (designated here): US01013695!; isolectotypes: GH!, NY!, US!). Trelease (1924, p. 205, 230) cited just one collection number, but he did not indicate where the specimen was deposited .

Quercus candicans auct. non Neé f. michoacana Trel., Mem. Natl. Acad. Sci. 20: 204, pl. 418(1) (1924). Type:— MEXICO, Michoacán: C. G. Pringle 3955 (Lectotype: US01013694! designated by Valencia-A. et al. (1918, p. 256); isolectotypes: A!, BM!, BR!, E!, F!, G!, HBG (image)!, K!, KFTA (image)!, M!, MEL (image)!, MEXU!, MICH!, MO!, MU (image)!, NY!, P!, PH!, S (image)!, UC!, VT!). Although Trelease designated Pringle 3955 as the type collection, we have not found any specimens that match the photograph in plate 418(1).

Quercus chimaltenangana Trel. f. gemmata C.H.Mull., Amer. Midl. Naturalist 18(5): 855 (1937). Type:— GUATEMALA, Quiche: A. F. Skutch 1654 (Holotype: ILL00007436 !; isotypes: A!, F!, NY!, UCSB (fragment)!, US!) .

Quercus flavida Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 178 (1854) . Type:— MEXICO, Puebla: F. M. Liebmann 3478 (Lectotype (designated here): C 10012466!; isolectotype: CAS!). Liebmann 3478 was represented at C by two specimens with sheet numbers of Cupulif. Libm. 1 and Cupulif. Libm. 2. On p. 203 Trelease (1924) wrote “ Liebmann, 2, 3478, May, 1841 the type of Q. flavida Liebmann ” and on p. 230 in the caption to pl. 412(2) he wrote “The type of Q. flavida (Mexico, Liebmann, 2). Photographed by permission at Copenhagen.” The image in pl. 412(2), however, Liebmann 3478 Cupulif. Lbm. 1 is now at CAS and Liebmann 3478 Cupulif. Lbm. 2 is at C. Because of this ambiguity, we are lectotypifying the C specimen.

Quercus intermedia M.Martens & Galeotti non Boenn. ex Rchb. (1831), Bull. Acad. Roy. Sci. Bruxelles 10(1): 223 (1843). See Q. calophylla f. intermedia above.

Quercus tonaguiae Trel., Mem. Natl. Acad. Sci. 20: 190, pl. 385 (1924). Type:— MEXICO, Oaxaca: H. Galeotti 105 (Holotype at B destroyed. Lectotype (designated here): BR0000030524842 !; isolectotypes: G!, K!, P!). The holotype at B was Galeotti s. n. Trelease (1924, p. 190) wrote that the specimen in Berlin was “referred to be Martens and Galeotti, p. 224, under no. 105, to Q. acutifolia .” For this reason we are lectotypifying the BR number as 105 which were the specimens referred to by M. Martens & Galeotti (1843, p. 224). The placement of this species needs additional study, but we consider the type to fall within the variation found within Q. calophylla as broadly interpreted.

Distribution: Guatemala, Mexico (Chiapas, Chihuahua, Ciudad de México, Colima, Durango, Guanajuato, Guerrero,

Hidalgo, Jalisco, México, Michoacán, Morelos, Nayarit, Oaxaca, Puebla, Querétaro, San Luis Potosí, Sinaloa, Sonora ,

Tamaulipas, Tlaxcala, Veracruz, Zacatecas). The occurrence of Q. calophylla in Tamaulipas reported by Villaseñor

(2016) needs to be confirmed.

Quercus canbyi Trel., Mem. Natl. Acad. Sci. 20: 188, pl. 379b (1924). Type :— MEXICO, Nuevo León: C. G. Pringle 2393 (Holotype at B destroyed. Lectotype (designated here): M-0244476!; isolectotypes: AC!, BM!, BR!, CAS!, E!, ENCB (image) !, F!, G!, GH!, HBG (image) !, K!, MEL (image)!, MEXU!, MICH!, MO!, NDG (image)!, NY!, P!, PH!, S (image) !, UC!, UCSB (fragment) !, US!, VT !).

Quercus alamarensis C.H.Mull., J. Arnold Arbor. 17(3): 174 (1936). Type:— MEXICO, Nuevo León: C. H. Muller & M. T. Muller 613 (Holotype: A00055614 !; isotypes: MICH!, UCSB (fragment)!).

Quercus canbyi f. ascendens Trel., Mem. Natl. Acad. Sci. 20: 188, pl. 380a (1924). Type:— MEXICO, Nuevo León: C. G. Pringle 11705 (Holotype at B destroyed. Lectotype (designated here): US00089467!; isolectotypes: A!, C!, CAS!, GH!, K!, L!, LE!, MICH!, MO!, L!, VT!).

Quercus canbyi f. attenuata C.H.Mull., J.Arnold Arbor. 17(3): 175 (1936). Type:— MEXICO, Nuevo León: C. H. Muller & M. T. Muller 1347 (Holotype: A00055638 !; isotypes: TEX (image)!, UCSB (fragment)!).

Quercus canbyi f. berlandieri Trel., Mem. Natl. Acad. Sci. 20: 188, pl. 380b (1924). Type:— MEXICO, Tamaulipas: J. Berlandier 2221 (Holotype: G; isotypes: CAS!, G!, GH!, K!, NY!, P!, UCSB (fragment)!). There are several sheets at G marked as isotypes, but none of them match the holotype shown in Trelease (1924) pl. 380b.

Quercus canbyi f. concolor Trel. & C.H.Mull., Bull. Torrey Bot. Club 63(3): 152 (1936). Type:— MEXICO, Nuevo León: C. H. Muller & M. T. Muller 590 (p.p.) (Holotype: ILL00007466 !; isotypes:TEX (image)!,UCSB (fragment)!). The specimen of C. H. Muller & M. T. Muller 590 at A is Q. glaucoides , so it is not an isotype.

Quercus canbyi f. karwinskii (Trel.) C.H.Mull., J. Arnold Arbor. 17(3): 175 (1936).

Quercus canbyi f. pedunculata C.H.Mull., J. Arnold Arbor. 17(3): 175 (1936). Type:— MEXICO, Nuevo León: C. H. Muller & M. T. Muller 1009 (Holotype: A00055641 !; isotype: UCSB (fragment)!).

Quercus canbyi f. setacea C.H.Mull., J. Arnold Arbor. 17(3): 175 (1936). Type:— MEXICO, Nuevo León: C. H. Muller & M. T. Muller 1011 (Holotype: A00055640 !; isotype: UCSB (fragment)!).

Quercus canbyi f. subovatifolia C.H.Mull., J. Arnold Arbor. 17(3): 175 (1936). Type:— MEXICO, Nuevo León: C. H. Muller & M. T. Muller 1346 (Lectotype (designated here): A00069937 !; isolectotype: UCSB (fragment)!). Muller (1936) gave this form a name and cited three collection numbers, but he did not designate a type.

Quercus karwinskii Trel., Mem. Natl. Acad. Sci. 20: 188, pl. 379a (1924). Type :— MEXICO: W. F. Karwinski s. n. (Holotype: M!). Trelease (1924. p. 189) wrote “Without data (Karwinski, the type in the Munich herbarium)” and on p. 228 for the caption to pl. 379b he wrote “The type (Mexico, Karwinski).” Plate 379b shows that the sheet at M has the number 464 written on the linen tape holding down the specimen. The tradition of writing numbers on the linen tape is seen on other M specimens and is not to be considered a collection number but rather a consecutive numbering done at M some time during the 19th century for specimens in each family (H.-J. Esser, pers. comm.).

Distribution: Mexico (Chihuahua, Coahuila, Hidalgo, Nuevo León, San Luis Potosí, Tamaulipas).

Quercus castanea Neé, Anales Ci. Nat. 3: 276 (1801). Type:— MEXICO, Hidalgo: L. Neé s. n. (1791) (Lectotype: MA25950 ! designated by Bartholomew & Almeda (2022, p. 28); isolectotypes: MA!, UCSB (as to larger leaf from MA lectotype).

Quercus axillaris E.Fourn. ex Trel., Mem. Natl. Acad. Sci. 20: 176, pl. 352a (1924). Type :— MEXICO, Ciudad de México: E. Bourgeau 1138 (Syntypes at B destroyed and at G. Lectotype (designated here): G (00410066) !; isolectotypes: BR!, C!, GH!, K!, MO (fragment)!, MPU!, P!, S (image) !, UCSB (image)!, US!). Trelease (1924, p. 227, pl. 352) cites the type as being both at B and G . There is a sheet at G marked as an isotype which is in fact an isosyntype as it does not match the syntype in Trelease (1924) pl. 352a. We are lectotypifying the name based on the existing isosyntype at G .

Quercus castanea var. sublobata A.DC., Prodr. 16(2.1): 72 (1864). Type :— GUATEMALA: J. R . R. Warszewicz 10 (Holotype: G!) .

Quercus circummontana Trel., Mem. Natl. Acad. Sci. 20: 177, pl. 356 (1924). Type :— MEXICO, Veracruz: F. M. Liebmann 3455 (Lectotype (designated here): C 10012544!; isolectotypes: C!, CAS!, MO!, NY!, UCSB (fragment)!). Of the three syntype sheets of 3455 originally at C , one is now at CAS. Trelease (1924, pl. 356) shows closeup images of two sheets of 3455 at C . The bottom image in plate 356 is selected as the lectotype, and the top image is the isolectotype at CAS .

Quercus consociata Trel., Mem. Natl. Acad. Sci. 20: 172, pl. 345a (1924). Type :— GUATEMALA: J. R. R. Warszewicz 9 (Holotype at B destroyed. Lectotype (designated here): JE00006775 !) .

Quercus crassivenosa Trel., Repert. Spec. Nov. Regni Veg. 33: 316 (1934). Type:— MEXICO, Guerrero: L. Schultze 33 (Holotype at B destroyed. Lectotype (designated here): ILL00007524 ! (fragment)).

Quercus impressa Trel., Mem. Natl. Acad. Sci. 20: 181, pl. 364a (1924). Type :— MEXICO, Oaxaca: C. G. Pringle 6029 (Holotype at B destroyed. Lectotype (designated here): P00754104 !; isolectotypes: A!, AC!, BM!, BR!, CAS!, E!, ENCB (image)!, G!, GH!, HBG (image) !, K!, KFTA (image)!, MEL (image)!, MEXU!, MO!, NDG (image)!, NY!, P!, PH!, S (image) !, UCSB (fragment) !, UC!, US!, VT !).

Quercus lanigera M.Martens & Galeotti, Bull.Acad. Roy. Sci. Bruxelles 10(1): 215 (1843). Type:— MEXICO, Oaxaca: H. Galeotti 104 (Lectotype: BR0000006912321 ! designated by Trelease (1924, p. 228, pl. 354); isolectotypes: BR!, G!, K!, P!, UCSB (fragment)!). Although M. Martens & Galeotti (1843) cite only one collection, there are two sheets of this number at BR marked as types, one of which one was effectively lectotypified by Trelease.

Quercus lanigera f. sideroxyloides Trel., Mem. Natl. Acad. Sci. 20: 177, pl. 355 (1924). Type :— MEXICO, Puebla: F. G . Nicolas s. n. (1912) (Holotype: F!) .

Quercus mucronata Willd., Sp. Pl. , ed. 4 4(1): 436 (1805). This is a superfluous name as the older name Q. castanea (1801) is erroneously cited as a synonym.

Quercus pulchella Bonpl., Pl. Aequinoct. 2: 44, pl. 88 (1809). Type:— MEXICO, Michoacán: A. Bonpland 4230 (Holotype: P00129710 !; isotypes: P (as s.n.)!, UCSB (fragment)!).

Quercus rossii Trel., Mem. Natl. Acad. Sci. 20: 179, pl. 362b (1924). Type:— MEXICO, Michoacán: H. Ross 374 (Holotype at B destroyed. Lectotype (designated here): M-0244474!; isolectotypes UCSB (fragment from M lectotype)!).

Quercus scherzeri Trel., Mem. Natl. Acad. Sci. 20: 172, pl. 343b (1924). Type :— HONDURAS, Comayagua: K. R. von Scherzer s. n. (Apr 1854) (Lectotype (designated here): image in Trelease (1924, pl. 343b)!). No extant original material exists, so we are lectotypifying the name based on the image of the holotype.

Quercus seleri Trel., Mem. Natl. Acad. Sci. 20: 181, pl. 364b (1924). Type:— MEXICO, Morelos: C. Seler & E. G. Seler 4302 (Holotype at B destroyed. Lectotype (designated here): image in Trelease (1924, pl. 364b)!). No extant original material exists, so we are lectotypifying the name based on the image of the holotype.

Quercus serrulata Trel., Mem. Natl. Acad. Sci. 20: 179, pl. 361a (1924). Type:— MEXICO, Jalisco: M. E. Jones 445 (Holotype: US 00089640!) .

Quercus spathulistipula Trel., Repert. Spec. Nov. Regni Veg. 33: 319 (1934). Type:— MEXICO, Guerrero: L. Schultze 332 (Holotype: at B destroyed). The determination of Q. spathulistipula as Q. castanea is based on the Trelease (1934) description because no images or original specimens exist.

Quercus subcrispata Trel., Mem. Natl. Acad. Sci. 20: 176, pl. 353 (1924). Type:— MEXICO, Puebla: G. Arsène 5926 (Holotype: US 00089662!; isotypes: BM!, ILL!, MPU!, PH!) .

Quercus tepoxuchilensis Trel., Mem. Natl. Acad. Sci. 20: 180, pl. 363a (1924). Type:— MEXICO, Puebla: G. Arsène 201 (Holotype at B destroyed. Lectotype (designated here): US 00089678!; isolectotypes: MEXU!, US!) .

Quercus tepoxuchilensis f. perplexans Trel., Mem. Natl.Acad. Sci. 20: 180 (1924). Type:— MEXICO, Puebla: G. Arsène 2068 (Holotype: ILL00007569 !; isotypes: MO!, MPU!, US!) .

Quercus tristis Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 174 (1854) . Type:— MEXICO, Veracruz: F. M. Liebmann 3455 (Lectotype (designated here): C 10012544!; isolectotypes: C!, CAS!, MO!, NY!). Trelease (1924, p. 227), in the caption to pl. 341, indicated two collections as the types, so this did not constitute lectotypification. The placement of this species needs additional study, but we consider the type to fall within the variation found within Q. castanea as broadly interpreted.

Quercus tristis f. mixcoensis Trel., Mem. Natl. Acad. Sci. 20: 172 (1924). Type :— GUATEMALA, Sacatepequez: W. Trelease 31 (Holotype: ILL00007553 !) .

Quercus tristis f. niederleini Trel., Mem.Natl.Acad.Sci. 20: 171, pl.343a (1924). Type :— EL SALVADOR: G. Niederlein s. n. (9 Jan 1898) (Holotype at B destroyed. Lectotype (designated here): image in Trelease (1924, pl. 343a)!). No extant original material exists, so we are lectotypifying the name based on the image of the holotype.

Quercus tristis f. sublobata (A.DC.) Trel., Mem. Natl. Acad. Sci. 20: 171 (1924).

Quercus tristis f. vulcani Trel., Mem. Natl. Acad. Sci. 20: 171, pl. 342b (1924). Type :— GUATEMALA, Escuintla: K. R. von Scherzer s. n. (Holotype at W destroyed. Lectotype (designated here): image in Trelease (1924, pl. 342b)!). No extant original material exists, so we are lectotypifying the name based on the image of the holotype.

Quercus verrucosirama Trel., Repert. Spec. Nov. Regni Veg. 33: 319 (1934). Type:— MEXICO, Guerrero: L. Schultze 382 (Holotype: at B destroyed). The determination of Q. verrucosirama as probably Q. castanea is based on the Trelease (1934) description as no images or original specimens exist.

Distribution: El Salvador, Guatemala, Honduras, Mexico (Aguascalientes, Chiapas, Chihuahua, Ciudad de México,

Coahuila, Colima, Durango, Guanajuato, Guerrero, Hidalgo, Jalisco, México, Michoacán, Morelos, Nayarit, Nuevo

León, Oaxaca, Puebla, Querétaro, San Luis Potosí, Sinaloa, Sonora, Tamaulipas, Tlaxcala, Veracruz, Zacatecas).

The occurrence of Q. castanea in Chihuahua, Colima, and Nuevo León reported by Villaseñor (2016) needs to be confirmed.

Quercus cedrosensis C.H.Mull., Madroño 16(6): 188, fig. 1–3, 5 (1962). Type :— MEXICO, Baja California Sur: C. H. Muller 10775 (Holotype: UCSB000021 !; isotypes: CAS!, G!, UCSB (fragment)!, US!, VT!) .

Distribution: Mexico (Baja California, Baja California Sur). Quercus chrysolepis Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 173 (1854) . Type:— U.S.A., California: T. Hartweg 1962 (Lectotype: K000832349! designated by Trelease (1924, p. 222, pl. 209); isolectotypes: K!, UCSB (fragment from K lectotype)!). Liebmann (1854, p. 173) cited six syntypes.

Distribution: Mexico (Baja California, Chihuahua, Sonora), U.S.A. (Arizona, California, Nevada, New Mexico,

Oregon).

Quercus coahuilensis Nixon & C.H.Mull., Brittonia 45(2): 150, fig. 2 (1993). Type :— MEXICO, Coahuila : I. M. Johnston & C. H. Muller 567 (Holotype: BH (image)!). This species is part of the Q. hypoxantha complex that could also be treated in a broad sense as a single species.

Distribution: Mexico (Chihuahua, Coahuila).

Quercus coffeicolor Trel., Mem. Natl. Acad. Sci. 20: 137, pl. 261a (1924). Type:— MEXICO, Sinaloa: J. N. Rose 1758 (Holotype: US00089482!; isotypes: GH!, K!, MO!, NY!, UCSB (fragment)!).

Quercus aequivenulosa Trel., Mem. Natl. Acad. Sci. 20: 138, pl. 261b (1924). Type:— MEXICO, Nayarit: J. N. Rose 2231 (Holotype: US00089387!; isotypes: GH!, NY!, UCSB (fragment)!).

Quercus mexiae L.M.González, Ibugana 9: 60, fig. 5–7 (2018). Type:— MEXICO, Jalisco: L.M.González V. & J.A. Pérez de la Rosa 5017 (Holotype: IBUG0177437). Although we have not seen the holotype, from the description and from those paratypes seen by Breedlove and us, we consider that Q. mexiae is not sufficiently different to justify it as a distinct species from Q. coffeicolor .

Quercus prainiana Trel. Non H. Lév. (1913) ,Mem. Natl.Acad. Sci. 20:138, pl. 262a, pl. 262b (1924). Type:— MEXICO, Jalisco: C. G. Pringle 8854 (Syntypes: C 10012523!, K000512914 !; isosyntypes: A!, BM!, CAS!, COLO (image)!, E!, ENCB (image)!, F!, G!, GH!, HBG (image)!, ILL!, L!, LL (image)!, MEXU!, MICH!, MO!, MSC (image)!, NY!, P!, PH!, UC!, UCSB (fragment)!, US!, VT!). Trelease (1924, p. 138) designated a type collection number, but on p. 224 in the caption to pl. 262 he did not specify which herbarium, C or K, was to house the holotype. In Plate 262 he has images of one specimen at C and one at K, both of which he called the type which makes both the C and K specimens syntypes. Because this species is a later homonym we are not lectotypifying the name.

Distribution: Mexico (Colima, Durango, Jalisco, Nayarit, Querétaro, San Luis Potosí, Sinaloa, Zacatecas).

Quercus confertifolia Bonpl., Pl. Aequinoct. 2: 53, pl. 94 (1809). Type:— MEXICO, Guanajuato: A. Bonpland 4407 (Holotype: P00129738 !; isotypes: P!, UCSB (fragment)!). Although lectotypified by Valencia-A. & Coombes (2020, p. 89), this was not necessary because the holotype at P has the specimen label in Bonpland’s hand. The isotype most likely was added to P or identified as such at a later date.

Quercus castanea Neé var. glabrata (Liebm. ex Seem.) A.DC., Prodr. 16(2.1): 72 (1864).

Quercus gentryi C.H.Mull., Amer. Midl. Naturalist 27(2): 474 (1942). Type :— MEXICO, Sinaloa: H. S. Gentry 5686 (Holotype: US 00089541!; isotypes: DS!, GH!, MEXU!, MICH!, MO!, NY!, RSA (image)!, UCSB (fragment from US holotype)!, UCSB (fragment)!). The holotype at US was formerly at NA .

Quercus mexicana Bonpl. f. confertifolia (Bonpl.) Trel., Mem. Natl. Acad. Sci. 20: 174 (1924).

Quercus mexicana f. glabrata Trel., Mem. Natl. Acad. Sci. 20: 174 (1924). Type:— MEXICO, Jalisco: H. Galeotti 118 (Lectotype (designated here): BR0000005105380 !; isolectotypes: BR!, G!, P!). We are lectotypifying one of the syntypes.

Quercus mexicana var. confertifolia (Bonpl.) Wenz., Jahrb. K ̂nigl. Bot. Gart. Berlin 3: 209 (1884).

Quercus mexicana var. glabrata Liebm. ex Seem., Bot. Voy. Herald 332 (1856). Type:— MEXICO, Sinaloa: B. C. Seemann 1974 (Holotype: K000512942 !; isotypes: BM!, C!, GH!). Valencia-A. & Coombes (2020, p. 90) designated Seemann 1974 at BM ( BM000643467 !) as the lectotype, but this is an invalid lectotypification, because the holotype as designated by Seemann is in the Hooker Herbarium at K and is the specimen with the herbarium name of Quercus mexicana var. glabrata in Liebmann’s hand.

Distribution: Mexico (Aguascalientes, Chihuahua, Colima, Durango, Guanajuato, Hidalgo, Jalisco, México,

Michoacán, Nayarit, Querétaro, Sinaloa, Zacatecas).

Quercus conzattii Trel., Proc. Amer. Philos. Soc. 60: 33, pl. 4 (1921). Type :— MEXICO, Oaxaca: C. Conzatti 1900 (Holotype: F0060021 F!; isotype: MEXU!) .

Quercus urbani Trel. f. parvifolia Martínez, Anales Inst. Biol. Univ. Nac. Autón. México 37(1 & 2): 93 (1966). This form is based on Q. conzattii , so the name has the same type.

Distribution: Mexico (Durango, Guerrero, Jalisco, Nayarit, Oaxaca, Zacatecas). Quercus cornelius-mulleri Nixon & K.P.Steele, Madroño 28(4): 210, fig. 4, 5 (1981). Type:— U.S.A., California: K. C. Nixon & K. P. Steele 2765 (Holotype: BH; isotypes: DAV, GH!, MO!, NY!, RSA (image)!, SD (image)!, TEX (image)!, UC!).

Distribution: Mexico (Baja California), U.S.A. (California).

Quercus corrugata Hook., Icon. Pl. 5: sub pl. 403, 404 (1842). Type:— GUATEMALA, San Marcos: G. V. Skinner 5 (Holotype: K000512592 !; isotypes: C (as s.n.?)!, K!, MPU!). The occurrence in Mexico by Villaseñor (2016) may actually be collections of Q. lancifolia Schltdl. & Cham. , but additional study is needed.

Quercus corrugata var. granulifera Trel., Mem. Natl. Acad. Sci. 20: 45, pl. 24 (1924). Type :— GUATEMALA: J. R. R. Warszewicz 11 (Holotype at B destroyed. Lectotype (designated here): G00410063 !; isolectotypes: K!, P!, UCSB (fragment)!).

Quercus corrugata var. ipalensis Trel., Mem. Natl. Acad. Sci. 20: 45 (1924). Type :— GUATEMALA, Chiquimula: H. Pittier 1869 (Holotype: US00089474!). The specimen at US is marked by Trelease and in his hand as “ Q. corrugata iipalensis Trel. ”

Quercus corrugata var. microcarpa Wenz., Jahrb. K ̂nigl. Bot. Gart. Berlin 3: 192 (1884). Type:— COSTA RICA, San José: Polakovsky s. n. (Holotype: at B destroyed). The determination is based on the Wenzig (1884) description as no images or original specimens exist.

Distribution: Costa Rica, Guatemala.

Quercus cortesii Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 175 (1854) . Type:— MEXICO, Veracruz: F. M. Liebmann 3463 (Lectotype: C10013674 ! designated by Trelease (1924, p. 228, pl. 373b); isolectotypes: G! UCSB (fragment from C lectotype)!).

Quercus anglohondurensis C.H.Mull., U.S.D.A. Bur. Pl. Industr. Misc. Publ. 477: 76, pl. 114, 115 (1942). Type:— BELIZE, El Cayo: C. L. Lundell 6615 (Holotype: MICH1109366 !; isotypes: DS!, GH!, MEXU!, MO!, NY!, TEX (image)!, UCSB (fragment)!, US!, WIS!).

Quercus brenesii Trel., Mem. Natl. Acad. Sci. 20: 186, pl. 377b (1924). Type:— COSTA RICA, Alajuela: A. Brenes 14520 (Holotype at B destroyed. Lectotype (designated here): US 00089436!; isolectotypes: GH!, K!, UCSB (fragment)!, US!) .

Quercus gracilior C.H.Mull., U.S.D.A. Bur. Pl. Industr. Misc. Publ. 477: 77, pl. 116 (1942). Type:— HONDURAS, Comayagua: J. B. Edwards 285 (Holotype: A00055575 !; isotypes: F!, UCSB (fragment)!).

Quercus gulielmi-treleasei C.H.Mull., U.S.D.A. Bur. Pl. Industr. Misc. Publ. 477: 58, pl. 79, 80 (1942). Type :— PANAMA, Chiriquí: R. J. Seibert 226 (Holotype: MO-143075!).

Quercus huitamalcana Trel., Contr. U.S. Natl. Herb. 23(2): 196 (1922). Type:— MEXICO, Puebla: F. M. Liebmann 3560 (Lectotype: C10013681 ! designated by Trelease (1924, p. 186); isolectotypes: C!, CAS!, UCSB (fragment from C lectotype)!). Nom. nov. for Q. serra Liebm. (1854) non Unger (1845). The lectotype is sheet Cupulif. Liebm. 31 of Liebmann 3560.

Quercus paxtalensis C.H.Mull., U.S.D.A. Bur. Pl. Industr. Misc. Publ. 477: 75, pl. 113 (1942). Type:— MEXICO, Chiapas: E. Matuda 1724 (Holotype: MICH1109345 !; isotypes: A!, K!, LL (image)!, MO!, NY!, US!, VT!).

Quercus serra Liebm. non Unger (1845), Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 174 (1854). See Q. huitamalcana above.

Quercus tenuiaristata Trel., Publ. Field Mus. Nat. Hist., Bot. Ser. 17(4): 358 (1938). Type:— HONDURAS, Comayagua: T. G. Yuncker et al. 6204 (Holotype: ILL00007574 !; isotypes: F!, G!, GH!, K!, MICH!, MO!, NY!, UCSB (fragment)!).

Distribution: Belize, Costa Rica, El Salvador, Honduras, Mexico (Chiapas, Colima, Guerrero, Jalisco, Oaxaca, Puebla,

Tamaulipas, Veracruz), Nicaragua, Panama.

Quercus costaricensis Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 184 (1854) . Type:— COSTA RICA, Cartago: A. S. Oersted 3465 (Lectotype (designated here): C 10012470!; isolectotypes: BM!, C!, CAS!, G!, UCSB (fragment)!). Trelease (1924, p. 225) in the caption to pl. 281 wrote “The type (Costa Rica, Oersted, 3465).” In pl. 281 he shows photos taken of the two syntypes now at C, so Trelease’s statement cannot be considered effective lectotypification. We are lectotypifying the specimen shown in the bottom photo of pl. 281.

Quercus endresii Trel., Mem. Natl. Acad. Sci. 20: 145, pl. 280 (1924). Type :— COSTA RICA: A. R. Endres s. n. (Holotype at W destroyed. Lectotype (designated here): image in Trelease (1924, pl. 280)!). No extant original material exists, so we are lectotypifying the name based on the image of the holotype.

Quercus irazuensis Kuntze, Revis. Gen. Pl. 2: 641 (1891). Type:— COSTA RICA, Cartago: O. Kuntze 2344 (Holotype: NY00253700 !; isotypes: K!, UCSB (fragment)!).

Quercus wesmaelii Trel., Mem. Natl. Acad. Sci. 20: 172, pl. 344a (1924). Type :— COSTA RICA, Alajuela: H. Pittier 773 (Holotype at B destroyed. Lectotype (designated here): BR0000006914523 !; isolectotype: BR!) .

Distribution: Costa Rica, Panama.

Quercus crassifolia Bonpl., Pl. Aequinoct. 2: 49, pl. 91 (1809). Type :— MEXICO, Guerrero: A. Bonpland 4142 (Holotype: P00129735 !; isotype: P!) .

Quercus brachystachys Benth., Pl. Hartw. 91 (1842). Type:— GUATEMALA, Guatemala: T. Hartweg 618 (Holotype: K000512503 !; isotypes: BM!, CGE!, K!, LD (image)!).

Quercus chicamolensis Trel., Contr. U.S. Natl. Herb. 23(2): 190. (1922). Type :— MEXICO, Oaxaca: H. Galeotti 102 (Holotype: BR0000005104734 !; isotypes: BR!, G!, P!, UCSB (fragment)!). Nom. nov. for Q. mollis M.Martens & Galeotti (1843) non Raf. (1838). Although Trelease (1924, p. 224) stated that the specimen in pl. 246 at BR was the type, this specimen is an isotype. The holotype is the specimen from M . Martens’ own herbarium now at BR .

Quercus crassifolia var. errans (Trel.) E.F.Warb., Bull. Misc. Inform. Kew 1939 (2): 90 (1939).

Quercus errans Trel., Mem. Natl. Acad. Sci. 20: 131, pl. 245a (1924). Type:— MEXICO, Hidalgo: T. Hartweg 424 (Holotype at B destroyed. Lectotype (designated here): G00177569!; isolectotypes: A!, CGE!, G!, K!, LE!, NY!, OXF!, P!). There is no extant collection of the holotype, Hahn 667, which was collected at Perote in Veracruz, Mexico. From the number sequence, Hahn 667 would have been collected in 1865. There is a specimen of Hahn s. n. (1 Jul 1866) both at K and P, but this collection cannot be considered an isotype, as it was collected a year later than the holotype, even though it was collected at the same locality and is an excellent match for the photo of the holotype in Trelease (1924), pl. 245a. On page Trelease (1924) cites several paratypes which include Ehrenberg 895, 896, 920, 1021, and 1284 which, although not stated, were probably all seen by him at B and were destroyed. Of these five collections, the only one that is extant is Ehrenberg 896 from Hidalgo, Mexico now at NY and VT. The photo in pl. 245b is of Hartweg 424 which Trelease also states is “a second collection” of Q. errans , so it is also a paratype. The photo of this Hartweg collection was also taken of the specimen at B which is now destroyed. We are choosing Hartweg 424 at G as the lectotype, which is also the lectotype of Q. splendens var. pallidior .

Quercus felipensis Trel., Mem. Natl. Acad. Sci. 20: 129, pl. 237 (1924). Type :— MEXICO, Oaxaca: C. L . Smith 776 (Holotype: MO-144156!; isotype: VT!) .

Quercus miguelitensis Trel., Mem. Natl. Acad. Sci. 20: 132, pl. 249b (1924). Type :— MEXICO, San Luis Potosí: J. G. Schaffner 897 (Holotype: GH00055673 !; isotype: K!) .

Quercus mollis M.Martens & Galeotti non Raf. (1838), Bull. Acad. Roy. Sci. Bruxelles 10(1): 216 (1843). See Q. chicamolensis above.

Quercus moreliana Trel., Mem. Natl. Acad. Sci. 20: 130, pl. 241 (1924). Type:— MEXICO, Michoacán: G. Arsène 6681 (Holotype: US 00089579!) .

Quercus orbiculata Trel., Mem. Natl.Acad. Sci. 20: 132, pl. 249a (1924). Type:— MEXICO, San Luis Potosí: C. C. Parry & E. Palmer 836 (Holotype: GH00062252 !; isotypes: E!, F!, K!, MO!, NY!, P!, PH!, UCSB (fragment)!, US!, WIS!).

Quercus spinulosa M.Martens & Galeotti, Bull. Acad. Roy. Sci. Bruxelles 10(1): 218 (1843). Type:— MEXICO, Veracruz: H. Galeotti 114 (Holotype: BR0000006916114 !; isotypes: G!, P!, UCSB (fragment)!).

Quercus splendens Neé var. pallidior A.DC., Prodr. 16(2.1): 57 (1864). Type:— MEXICO, Hidalgo: T. Hartweg 424 (Lectotype: G00177569 ! designated by Trelease (1924, p. 224, pl. 243a); isolectotypes: A!, CGE!, G!, K!, LE!, NY!, OXF!, P!). A. de Candolle (1864, p. 57) cited two syntypes.

Quercus stipularis Bonpl., Pl. Aequinoct. 2: 47, pl. 90 (1809). Type:— MEXICO, Veracruz: A. Bonpland s. n. (Holotype: P00129748 !; isotypes: P!, UCSB (fragment)!).

Distribution: Guatemala, Honduras, Mexico (Aguascalientes, Chiapas, Chihuahua, Ciudad de México, Coahuila,

Colima, Durango, Guanajuato, Guerrero, Hidalgo, Jalisco, México, Michoacán, Morelos, Nayarit, Nuevo León, Oaxaca ,

Puebla, Querétaro, San Luis Potosí, Sinaloa, Sonora, Tamaulipas, Tlaxcala, Veracruz, Zacatecas). The occurrence of Q.

crassifolia in Coahuila, Colima, Sinaloa, and Tamaulipas reported by Villaseñor (2016) needs to be confirmed.

Quercus crassipes Bonpl., Pl. Aequinoct. 2: 37, pl. 83 (1809). Type :— MEXICO, Guanajuato: A. Bonpland s. n. (Holotype: P00129734 !; isotypes: HAL!, P!) .

Quercus colimae Trel., Mem. Natl. Acad. Sci. 20: 174, pl. 349a (1924). Type:— MEXICO, Jalisco: Burnett s. n. (1913) (Holotype: USFS). We have been unable to find the holotype. The synonomy is based on Trelease (1924) pl. 349a and the description.

Quercus colimae var. zauzillo Trel., Mem. Natl. Acad. Sci. 20: 175, pl. 349b (1924). Type:— MEXICO, Jalisco: Burnett s. n. (1913) (Holotype: USFS). We have been unable to find the holotype. The synonomy is based on Trelease (1924) pl. 349b and the description.

Quercus crassipes var. angustifolia Bonpl., Pl. Aequinoct. 2: 38, pl. 84 (1809). Type:— MEXICO, Guanajuato: A. Bonpland s. n. (Holotype: P00129729 !; isotypes: HAL!, P!, UCSB (fragment)!).

Quercus cuajimalpana Trel., Mem. Natl. Acad. Sci. 20: 165, pl 324 (1924). Type:— MEXICO, Ciudad de México: G. Arsène 8866 (Holotype: US00089472!; isotypes: A!, F!, GH!, ILL!, L!, MO!, MPU!, NY!, UCSB (fragment)!).

Quercus malifolia Trel., Mem. Natl. Acad. Sci. 20: 175, pl. 348b (1924). Type :— MEXICO, Veracruz: H. Ross 1256 (Holotype at B destroyed. Lectotype (designated here): M-0244472!; isolectotype UCSB (fragment from M lectotype)!). It is likely that the holotype at B was actually Ross 1256 rather than Ross 256 which was also the conclusion of C . H . Muller who removed some fragments of the M specimens that are now at UCSB.

Quercus mexicana Bonpl. f. angustifolia (Bonpl.) Trel., Mem. Natl. Acad. Sci. 20: 174, pl. 345 (1924).

Quercus mexicana f. lanosa Trel., Mem. Natl. Acad. Sci. 20: 174, pl. 347a (1924). Type:— MEXICO, Michoacán: T. Hartweg 431 (Holotype at B destroyed. Lectotype (designated here): K000512959 !; isolectotypes: BM!, CGE!, E!, G!, K!, L!, LE!, NY!, OXF!, P!, UCSB (fragment)!).

Quercus mexicana f. perfertilis Trel., Mem. Natl. Acad. Sci. 20: 174 (1924). Type:— MEXICO, Veracruz: J. N. Rose & R. Hay 5675 (Lectotype (designated here): US00089580!; isolectotype: A!). Trelease (1924) did not specify the place of deposit for the type collections.

Quercus obovalifolia E.Fourn. ex Trel., Mem. Natl. Acad. Sci. 20: 175, pl. 351b (1924). Type:— MEXICO, Ciudad de México: E. Bourgeau 1014 (Holotype: C 10012505!; isotypes: BM!, G!, K!, P!, UCSB (fragment from C holotype)!, UCSB (fragment)!).

Distribution: Mexico (Aguascalientes, Chiapas, Ciudad de México, Coahuila, Colima, Durango, Guanajuato,

Guerrero, Hidalgo, Jalisco, México , Michoacán, Morelos, Nayarit, Nuevo León, Oaxaca, Puebla, Querétaro, San Luis

Potosí, Tamaulipas, Tlaxcala, Veracruz, Zacatecas).

Quercus crispipilis Trel., Mem. Natl. Acad. Sci. 20: 184, pl. 370b (1924). Type:— GUATEMALA, Huehuetenango: C. Seler & E. G. Seler 2667 (Holotype at B destroyed. Lectotype (designated here): GH00069875 !; isolectotypes: NY!, UCSB (fragment from GH lectotype)!, UCSB (fragment)!).

Quercus cerifera Trel., Mem. Natl. Acad. Sci. 20: 184, pl. 370a (1924). Type:— MEXICO, Chiapas: O. F. Cook 116 (Holotype: US 00089459!) .

Quercus cinnamomea Trel., Mem. Natl. Acad. Sci. 20: 184, pl. 371 (1924). Type:— GUATEMALA, Totonicapan: O. F. Cook 27 (Holotype: US00089485!). The protologue says Cook 27 which Breedlove recorded as seeing at US, but the US database says Cook 127 and indicates it is an isotype. Unfortunately, there is no image available for the US specimen to compare to the image of the holotype published by Trelease (1924, pl. 371).

Distribution: Guatemala, Mexico (Chiapas).

Quercus cualensis L.M.González, Brittonia 55(1): 49, figs. 1–3, 5 (2003). Type:— MEXICO, Jalisco: L. M. González V. et al. 3076 (Holotype: IBUG0070352 (image)!; isotypes: CAS!, F (image)!, IEB, MEXU, MICH (image)!, NMC, NY (image)!, TEX (image)!, UC, US (image)!, WIS (image)!, ZEA). This narrowly endemic species is similar to the more widespread species Q. iltisii , but there are a number of distinct character states presented in the protologue which justify its recognition.

Distribution: Mexico (Jalisco).

Quercus delgadoana S.Valencia, Nixon & L.M.Kelly, Novon 21(2): 275, fig. 1 (2011). Type:— MEXICO, Hidalgo: S. Valencia A. et al. 3748 (Holotype: FCME; isotypes: BH, HUAP, IBUG (image)!, K, MEXU (image)!, MO, NY). The illustration in Valencia et al. (2011, Fig. 1 ) and the isotype image seen from MEXU appear to be similar to Q. perseifolia , a species not mentioned in the protologue. This possible synonymy needs additional study.

Distribution: Mexico (Chiapas, Hidalgo, Oaxaca, Puebla, Querétaro, San Luis Potosí, Tabasco, Veracruz).

Quercus depressa Bonpl., Pl.Aequinoct. 2: 50, pl. 92 (1809). Type:— MEXICO, Hidalgo: A. Bonpland 4145 (Holotype: P00129727 !; isotypes: B-W (as s.n.)!, CGE (as s.n.)!, HAL!, P!, UCBG (fragment)!).

Quercus carnerosana Trel., Mem. Natl. Acad. Sci. 20: 183, pl. 369b (1924). Type :— MEXICO, Coahuila: C. G. Pringle 2802 (Holotype: A00055603 !; isotypes: GH!, MEXU!) .

Quercus charcasana Trel. ex A.Camus, Chênes Texte 3: 216, 1269 (1953). Type:— MEXICO, San Luis Potosí: C. L. Lundell 5382 (Lectotype (designated here): ILL00007535 !; isolectotypes: A!, DS!, K!, S (image)!, US!). Camus did not specify where the type collection was deposited. The placement of this species needs additional study, but we consider the type to fall within the variation found within Q. depressa as broadly interpreted.

Quercus subavenia Trel., Mem. Natl. Acad. Sci. 20: 169, pl. 338b (1924). Type:— MEXICO, Puebla: C. G. Pringle 8908 (Holotype: G!; isotypes: A!, BM!, C!, CAS!, E!, F!, FI!, GH!, HBG (image)!, ILL!, K!, L!, LL (image)!, M!, MO!, NY!, P!, PH!, UC!, UCSB (fragment)!, US!, VT!).

Distribution: Mexico (Ciudad de México, Coahuila, Hidalgo, México, Nuevo León, Oaxaca, Puebla, San Luis Potosí,

Tamaulipas, Veracruz, Zacatecas). The occurrence of Q. depressa in Ciudad de México and México reported by

Villaseñor (2016) needs to be confirmed.

Quercus depressipes Trel., Mem. Natl. Acad. Sci. 20: 90, pl. 144a (1924). Type :— MEXICO. Durango: P. Tenorio L. & C . Romero de T . 1196 Buenos Aires, 36 km al W de Tepehuanes, por la Brecha a topia, Mpio. Tepehuanes , 22 Jul 1982 (Holotype at B destroyed. Neotype (designated here): MEXU00345812 !; isoneotype: CAS!) ..

Quercus bocoynensis C.H.Mull., Amer. Midl. Naturalist 19(3): 584 (1938). Type:— MEXICO, Chihuahua: F. Shreve 8022 (Holotype: BH; isotypes: UCSB (fragment)!, US!). An isotype in the protologue is also cited as being at the Desert Laboratory, Tucson , Arizona .

Quercus oblongifolia Torr. var. pallidinervis Trel., Mem. Natl. Acad. Sci. 20: 91, pl. 146b (1924). Type :— MEXICO, Chihuahua: E. W. Nelson 6309 (Holotype: US 00169742!; isotypes: GH!, K!) .

Distribution: Mexico (Chihuahua, Durango, Guanajuato, Hidalgo, Jalisco, Nuevo León, Querétaro, San Luis Potosí,

Sinaloa, Sonora, Tamaulipas, Zacatecas), U.S.A. (Texas).

Quercus devia Goldman, Contr. U.S. Natl. Herb. 16: 322 (1916). Type :— MEXICO, Baja California Sur: E. W. Nelson & E. A . Goldman 7454 (Holotype: US 00089501!; isotype: A!) .

Quercus viminea Trel., Mem. Natl. Acad. Sci. 20: 123, pl. 222a (1924). Type:— MEXICO, Sonora: R. Hartman 342 (Holotype at B destroyed. Lectotype (designated here): US00898088!; isolectotypes: CAS!, GH!, K!, NY!). The holotype was Endlich 1280 from Chihuahua, Mexico, and no isotypes have been found. The lectotype is selected from one of the paratypes from Sonora, Mexico. Although Villaseñor (2016) considered this species to be distinct, we consider it to fall within Q. devia as broadly interpreted.

Distribution: Mexico (Aguascalientes, Baja California, Baja California Sur, Chihuahua, Durango, Guanajuato, Jalisco,

Nayarit, San Luis Potosí, Sinaloa, Sonora, Zacatecas), U.S.A. (Arizona). The occurrence of Q. devia in Aguascalientes reported by Villaseñor (2016) needs to be confirmed.

Quercus diversifolia Neé, Anales Ci. Nat. 3: 270 (1801). Type:— MEXICO, Guerrero: L. Neé s. n. (1791) (Lectotype: MA26468 ! designated by Bartholomew & Almeda (2022, p. 28); isolectotype: UCSB (fragment from MA lectotype)!).

Quercus centralis Trel., Mem.Natl.Acad.Sci. 20: 61, pl. 60a (1924).Type:— MEXICO, Ciudad de México: C. G. Pringle 10394 (Holotype at B destroyed. Lectotype (designated here): CAS0655142 !; isolectotypes: GH!, LE!, UC!, US!, VT!, WIS!). The holotype was Endlich 1365c, which was probably collected in Ciudad de México , but no isotypes are extant. We are lectotypifying the name based on a paratype also from Ciudad de México .

Quercus sanchez-colinii Martínez, Anales Inst. Biol. Univ. Nac. Autón. México 24(2): 269, fig. 15 (1953[1954]). Type :— MEXICO, México: E. Matuda 28760 (Holotype: MEXU 01231768!; isotypes: CAS!, MEXU!, UCSB (fragment)!).

Quercus texcocana Trel., Mem. Natl. Acad. Sci. 20: 81, pl. 117a (1924). Type:— MEXICO, México: R. Endlich 653 (Holotype at B destroyed. Lectotype (designated here): image in Trelease (1924, pl. 117a)!). No extant original material exists, so we are lectotypifying the name based on the image of the holotype. The placement of this species needs additional study, but we consider the type to fall within the variation found within Q. diversifolia as broadly interpreted.

Quercus texcocana var. ampla Trel., Mem. Natl. Acad. Sci. 20: 81, pl. 117b (1924). Type:— MEXICO, México: R. Endlich 655 (Holotype at B destroyed. Lectotype (designated here): image in Trelease (1924, pl. 117b)!). No extant original material exists, so we are lectotypifying the name based on the image of the holotype.

Quercus tomentosa Willd. nom. illigit. var. diversifolia (Neé) A.DC., Prodr. 16(2.1): 33 (1864).

Distribution: Mexico (Ciudad de México, Coahuila, Guanajuato, Guerrero, Hidalgo, México, Michoacán, Morelos, Nuevo León, Oaxaca, Puebla, Querétaro, San Luis Potosí, Tamaulipas, Tlaxcala, Veracruz). The occurrence of Q. diversifolia in Coahuila, Nuevo León, and Tamaulipas reported by Villaseñor (2016) needs to be confirmed.

Quercus dumosa Nutt., N. Amer. Sylv. 1(1): 7 (1842). Type:— U.S.A., California: T. Nuttall s. n. (May 1819) (Holotype: BM000092132 !; isotype UCSB (fragment from BM holotype)!).

Quercus dumosa f. diversifolia Trel., Mem. Natl.Acad.Sci. 20: 117, pl. 205a (1924). Type :— MEXICO, Baja California: T. Brandegee s. n. (21 Apr 1893) (Holotype: A00055679 !) .

Quercus dumosa f. microcarpa Trel., Mem. Natl. Acad. Sci. 20: 117, pl. 203b (1924). Type :— MEXICO, Baja California: E. Palmer 640 (Holotype: MO) .

Quercus dumosa f. populifolia Trel., Mem. Natl.Acad. Sci. 20: 117, pl. 204b (1924). Type :— MEXICO, Baja California: T. Brandegee s. n. (6 May 1893) (Holotype: A) .

Distribution: Mexico (Baja California, Chihuahua, Jalisco, Michoacán), U.S.A. (California).

Quercus durifolia Seemen, Bot. Jahrb. Syst. 29(1): 95 (1900). Type:— MEXICO, Durango: E. Palmer 771 (Holotype at B destroyed. Lectotype (designated here): US00089491!; isolectotypes: A!, BM!, CAS!, F!, GH!, K!, MO!, NY!, UC!, UCSB (fragment)!).

Quercus flocculenta C.H.Mull., J. Arnold Arbor. 17(3): 172 (1936). Type:— MEXICO, Nuevo León: C. H. Muller & M. T. Muller 1277 (Holotype: A00055663 !; isotypes: TEX (image)!, UCSB (fragment)!). The placement of this species needs additional study, but we consider the type to fall within the variation found within Q. durifolia as broadly interpreted.

Quercus flocculenta f. incisa C.H.Mull., J.Arnold Arbor. 17(3): 173 (1936). Type:— MEXICO, Nuevo León: C. H. Muller & M. T. Muller 1276 (Holotype: A00055664 !; isotype: UCSB (fragment)!).

Quercus flocculenta f. oblongifolia C.H.Mull., J. Arnold Arbor. 17(3): 173 (1936). Type:— MEXICO, Nuevo León: C. H. Muller & M. T. Muller 1229 (Holotype: A00055665 !; isotypes: BM!, G!, MEXU!, NY!, P!, S (image)!, US!) .

Quercus sipuraca Trel., Mem. Natl. Acad. Sci. 20: 182, pl. 366b (1924). Type:— MEXICO, Chihuahua: R. Endlich 1286 (Holotype at B destroyed. Lectotype (designated here): image in Trelease (1924, pl. 366b)!). No extant original material exists, so we are lectotypifying the name based on the image of the holotype.

Distribution: Mexico (Chihuahua, Coahuila, Durango, Nuevo León, Oaxaca, Querétaro, San Luis Potosí, Sinaloa,

Sonora, Tamaulipas, Zacatecas).

Quercus dysophylla Benth., Pl.Hartw. 55(1840).Type:— MEXICO, Veracruz: T. Hartweg421 (Holotype:K000512275!; isotypes: BM!, CGE!, E!, LD (image)!, OXF (as mislabel #1963)!, UCSB (fragment)!). This species has been considered by some to be of hybrid origin between Q. crassifolia and Q. crassipes (Govaerts & Frodin 1998, Villaseñor 2016, López C. et al. 2017), but we consider the type collection to not be a hybrid.

Quercus esperanzae Trel., Mem. Natl. Acad. Sci. 20: 132, pl. 248 (1924). Type :— MEXICO, Puebla: C. A. Purpus 5332 (Holotype: F0060028 F!; isotypes: BM!, GM!, BM!, MEXU!, MO!, UC!, UCSB (fragment)!). Although Trelease (1924, p. 224) in the caption for pl. 248 wrote “The type (Mexico, Purpus 5332). Photographed by permission at N. Y.” the actual specimen in plate 248 is at F .

Quercus floccosa Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 178 (1854) . Type:— MEXICO, Veracruz: F. M. Liebmann 3479 (Lectotype: C 10012481! Designated by Trelease (1924, p. 223, pl. 232a); isolectotypes: CAS! UCSB (fragment from C lectotype)!). The placement of this species needs additional study, but we consider the type to fall within the variation found within Q. dysophylla as broadly interpreted.

Quercus fournieri Trel., Mem. Natl. Acad. Sci. 20: 177, pl. 357 (1924). Type:— MEXICO, Veracruz: L. Hahn s. n. (Jun 1866) (Holotype: P00077457!; isotypes: BM (as 255)!, K (as 255)!, UCSB (fragment from P holotype)!). At P there are three sheets with the name Q. ferruginea , two of which have the number 258 and one which is s.n. This name was apparently being proposed by Fournier as a new species as “n sp.” is written on the s.n. sheet. Trelease (1924, p. 178) realized that this would be a later homonym of Q. ferruginea Michaux (1812) and published the new name, Q. fournieri , (Trelease 1924, p. 178, pl. 357). He cites “ Hahn 255, June, 1866” at P as the type although there is no Quercus specimen at P with the number Hahn 255. In the caption to plate 357 Trelease states “The type (Mexico, Hahn, 255)” which is in fact a close-up of the s.n. specimen at P. On the label for the P specimen of Hahn s. n. the date was originally written as “ Juillet 1866 ” with the month crossed out and in the same hand changed to “Juin.” Both BM and K have specimens distributed from P which are labeled Hahn 255 and dated as being collected in July 1866. With the mixup of numbers and dates but clearly Hahn s. n. at P (1924, pl. 357) being indicated as “the type”, the P specimen of Hahn s. n. at P is the holotype and the specimens of Hahn 255 at BM and K are probably isotypes. There is no mixup in the identity of any of these specimens as they are all Q. dysophylla . The type fragment at UCSB taken by C.H. Muller from a specimen at P is labeled “ Hahn 255?” indicating that Muller was aware of this number mixup by Trelease and also that P did not in fact have a collection of Hahn 255. The UCSB fragment which consists of a single leaf is undoubtedly a fragment from the Hahn s. n. holotype.

Quercus hahnii Trel., Mem. Natl. Acad. Sci. 20: 131, pl. 247 (1924). Type:— MEXICO, Ciudad de México: L. Hahn 347 (Holotype: C 10012474!; isotypes: P!, UCSB (fragment from C holotype)!).

Quercus imbricariifolia Trel., Mem. Natl. Acad. Sci. 20: 174, pl. 348ª (1924). Type:— MEXICO, Ciudad de México: E. Bourgeau 425 (Lectotype (designated here): P06814111!; isolectotypes: BM!, K!, MPU!, UCSB (fragment)!). The holotype, Endlich 1012, was destroyed at B; no isotypes have been found in any herbarium. We are lectotypifying the name based on the paratype. One of the specimens at P of Bourgeau 425 is Q. crassipes so it cannot be considered an isolectotype, but alternative interpretation would place this species as a synonym of Q. crassipes .

Quercus sagata E.F.Warb., Bull. Misc. Inform. Kew 1939 (2): 90 (1939). Type:— MEXICO, México: G. B. Hinton 6577 (Holotype: K000575022 !; isotypes: A!, F!, GBH (image)!, UCSB (fragment)!, US!). The status of this taxon needs additional study as the type has some similarities to Q. hintonii .

Distribution: Mexico (Chiapas, Ciudad de México, Durango, Guanajuato, Guerrero, Hidalgo, Jalisco, México,

Michoacán, Morelos, Nuevo León, Oaxaca, Puebla, Querétaro, San Luis Potosí, Tamaulipas, Tlaxcala, Veracruz,

Zacatecas).

Quercus eduardi Trel., Contr.U.S. Natl. Herb. 23(2): 189 (1922).Type:— MEXICO, Durango: E. Palmer 956 (Holotype at B destroyed. Lectotype (designated here): US 00089572!; isolectotypes: A!, CAS!, DS!, GH!, K!, MO!, NY!, UC!, UCSB!, US!). Nom. nov. for Q. oligodonta Seemen ex Loes. (1900) non Saporta (1879). This species is close to Q. laurina , and additional study is needed to determine if it is really distinct.

Quercus eduardi f. cespitifera Trel., Mem. Natl. Acad. Sci. 20: 122, pl. 219b (1924). Type:— MEXICO, Guanajuato: W. Schumann 1311 (Holotype at B destroyed. Lectotype (designated here): image in Trelease (1924, pl. 219a)!). No extant original material exists, so we are lectotypifying the name based on the image of the holotype.

Quercus oligodonta Seemen non Saporta (1879), Bot. Jahrb. Syst. 29(1): 96 (1900). See Q. eduardi above.

Distribution: Mexico (Durango, Guanajuato).

Quercus elliptica Neé, Anales Ci. Nat. 3: 278 (1801). Type:— MEXICO, Hidalgo: L. Neé s. n. (1791) (Lectotype: MA25956 ! designated by Bartholomew & Almeda (2022, p. 28); isolectotypes: MA!, UCSB (fragment from MA lectotype)!, UCSB (fragment)!). Although the type is reported to be from Hidalgo, no subsequent collections have been made from there. It is possible that Neé’s record was not correctly reported.

Quercus acutangula Trel., Repert. Spec. Nov. Regni Veg. 33: 318 (1934). Type:— MEXICO, Guerrero: L. Schultze 483 (Holotype: at B destroyed). The tentative determination of Q. acutangula as probably Q. elliptica is based on the Trelease (1934) description as no images, isotype, or other original material exist. Trelease, however, does mention the twigs being tomentose which is a characteristic of Q. elliptica , so the synonymy is uncertain.

Quercus atrescentirhachis Trel., Repert. Spec. Nov. Regni Veg. 33: 317 (1934). Type:— MEXICO, Guerrero: L. Schultze 347 (Syntypes at B destroyed. Lectotype (designated here): ILL00007471 ! (fragment)). ILL also has a fragment from Schulze 384 which Trelease (1934, p. 317) cites as “Also? (no. 384), with rather shallow cupules, 10–15 mm in diameter with apressed somewhat keeled scales and round-ovoid acorns.” Both L. Schultze 447 and 384 are Q. elliptica .

Quercus botryocarpa Trel., Mem. Natl. Acad. Sci. 20: 150, pl. 295 (1924). Type:— MEXICO, Nayarit: J. N. Rose 1971 (Holotype: US00089443!; isotypes: GH!, MO!, NY!, UCSB (fragment!)).

Quercus castanea Neé var. elliptica (Neé) Trel., Mem. Natl. Acad. Sci. 20: 178, pl. 359a (1924).

Quercus chiquihuitillonis Trel., Mem. Natl. Acad. Sci. 20: 139, pl. 264a (1924). Type:— MEXICO, Michoacán: E. Langlassé 212 (Holotype at B destroyed. Lectotype (designated here): P00754084 !; isolectotypes: G!, GH!, K!, MPU!, P!, UCSB (fragment)!, US!) .

Quercus coccinata Trel., Repert. Spec. Nov. Regni Veg. 33: 317 (1934). Type:— MEXICO, Guerrero: L. Schultze 305 (Holotype at B destroyed. Lectotype (designated here): ILL00007526 ! (fragment)).

Quercus comayaguana Trel., J. Arnold Arbor. 11(1): 25 (1930). Type:— HONDURAS, Comayagua: P. C. Standley 56229 (Holotype: F (accn. 581423)!; isotypes: A!, G!, ILL!, US!) .

Quercus exaristata Trel., Mem. Natl. Acad. Sci. 20: 140, pl. 266a (1924). Type:— MEXICO, Nayarit: J. N. Rose s. n. (3 Aug 1897) (Holotype: US 00089516!) .

Quercus guayabalana Trel., Publ. Field Mus. Nat. Hist., Bot. Ser. 8(1): 6 (1930). Type :— EL SALVADOR, Cuscatlán: S. Calderon 1982 (Holotype: US 00089528!; isotypes: GH!, ILL!, MO!) .

Quercus hondurensis Trel., Mem. Natl. Acad. Sci. 20: 140, pl. 266b (1924). Type:— HONDURAS, Cortés: C. Thieme 5440 (Holotype: US 00089556!) .

Quercus ignaciensis C.H.Mull., Amer. Midl. Naturalist 27(2): 475 (1942). Type:— MEXICO, Sonora: H. T. Green s. n. (26 Feb 1938) (Holotype: PH00023359!; isotype UCSB (fragment from PH holotype)!). The placement of this species needs additional study, but we consider the type to fall within the variation found within Q. elliptica as broadly interpreted.

Quercus incrassata Trel., Mem. Natl. Acad. Sci. 20: 147, pl. 287 (1924). Type:— MEXICO, Chiapas: E. Reeves 7 (Holotype: ILL00009824 !). The placement of this species needs additional study, but we consider the type to fall within the variation found within Q. elliptica as broadly interpreted.

Quercus langlassei Trel., Mem. Natl. Acad. Sci. 20: 138, pl. 263 (1924). Type:— MEXICO, Guerrero: E. Langlassé 1006 (Holotype at B destroyed. Lectotype (designated here): K000512915 !; isolectotypes: G!, GH!, P!, US!, UCSB (fragment)!).

Quercus linguifolia Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 180 (1854) . Type:— MEXICO, Oaxaca: F. M. Liebmann 3506 (Lectotype: C10014103 ! designated by Trelease (1924, p. 225, pl. 290); isolectotypes: CAS!, GH!, K (as s.n.)!, UCSB (from C lectotype)!). The K s.n. sheet is almost certainly a syntype as it is the right taxon from the right locality on the right date.

Quercus nectandrifolia Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 175 (1854) . Type:— MEXICO, Oaxaca: F. M. Liebmann 3509 (Lectotype: C10014106 ! designated by Trelease (1924, p. 225 pl. 289a); isolectotypes: CAS!, UCSB (fragment from C lectotype)!). The lectotype is sheet Cupulif. Liebm. 100.

Quercus oajacana Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 178 (1854) . Type:— MEXICO, Oaxaca: H. Galeotti 108 (Holotype: K000512934 !; isotypes: BR!, G!, P!, UCSB (fragment)!).

Quercus peradifolia E.F.Warb., Bull. Misc. Inform. Kew 1939 (2): 92 (1939). Type:— MEXICO, México: G. B. Hinton 6062 (Holotype: K000575017 !; isotypes: A!, BM!, CAS!, F!, G!, GBH (image)!, K!, NY!, UCSB (fragment)!).

Quercus porriginosa Trel., Repert. Spec. Nov. Regni Veg. 33: 316 (1934). Types: MEXICO, Guerrero. L. Schultze 330 and 333 (Syntypes: at B destroyed). No original material exists, so the only information is Trelease’s (1934) description. Quercus porriginosa to be a synonym of Q. elliptica , but Breedlove thought that it might be Q. peduncularis .

Quercus pubinervis M.Martens & Galeotti, Bull. Acad. Roy. Sci. Bruxelles 10(1): 211 (1843). Type:— MEXICO, Veracruz: H. Galeotti 134 (Holotype: BR!; isotypes: G!, P!, UCSB (fragment from BR holotype)!).

Quercus salicifolia Neé var. oajacana (Liebm.) Wenz., Jahrb. K ̂nigl. Bot. Gart. Berlin 3: 207 (1884).

Quercus yoroensis Trel., Publ. Field Mus. Nat. Hist., Bot. Ser. 9(4): 282 (1940). Type:— HONDURAS, Lempira: T. G. Yuncker et al. 8181 (Holotype: ILL00007541 !; isotypes: BM!, F!, G!, GH!, K!, MO!, NY!, S (image)!, UCSB (fragment)!, US!) .

Quercus yoroensis var. aguanana Trel., Publ. Field Mus. Nat. Hist., Bot. Ser. 9(4): 282 (1940). Type:— HONDURAS, Yoro: T. G. Yuncker et al. 8185 (Holotype: ILL00007540 !; isotypes: BM!, F!, G!, GH!, K!, MO!, NY!, S (image)!, UCSB (fragment)!, US!) .

Distribution: Belize, El Salvador, Guatemala, Honduras, Mexico (Chiapas, Chihuahua, Colima, Durango, Guerrero,

Hidalgo, Jalisco, México , Michoacán, Nayarit, Oaxaca, Puebla, Querétaro, San Luis Potosí, Sinaloa, Veracruz),

Nicaragua.Although the type of Q. elliptica is from Hidalgo, no subsequent collections of this species have been found in this state. The occurrence of Q. elliptica in Querétaro reported by Villaseñor (2016) also needs to be confirmed.

Quercus emoryi Torr. , Notes Milit. Reconn. 152, pl. 9 (1848). Type:— U.S.A., Texas: W. H. Emory s. n. (17 Oct 1846) (Lectotype: NY00248636 ! designated by Landrum (1992, p. 240)).

Quercus balsequillana Trel., Mem. Natl. Acad. Sci. 20: 123, pl. 220a (1924). Type:— MEXICO, Chihuahua: R. Endlich 1278 (Holotype at B destroyed. Lectotype (designated here): image in Trelease (1924, pl. 220a)!). Trelease (1924, p. 123) wrote that the type is Endlich 1275 but on p. 223 in the caption to pl. 220a he says the type is Endlich 1278. One or the other is probably a typographic error, but in any case, the type at B was destroyed.

Quercus duraznillo Trel., Mem. Natl. Acad. Sci. 20: 122, pl. 220b (1924). Type:— MEXICO, Chihuahua: C. G. Pringle 2112 (Holotype at B destroyed. Lectotype (designated here): A00970050!). The holotype was Endlich 1277b, and there are no extant isotypes. The lectotype is chosen from the only extant paratype which has spiny leaves like Q. duraznillo f. pinetorum , but both are Q. emoryi .

Quercus duraznillo f. bullata Trel., Mem. Natl. Acad. Sci. 20: 123, pl. 221a (1924). Type:— MEXICO, Chihuahua: R. Endlich 1277a (Holotype at B destroyed. Lectotype (designated here): image in Trelease (1924, pl. 221a)!). No extant original material exists, so we are lectotypifying the name based on the image of the holotype.

Quercus duraznillo f. cochutensis Trel., Mem. Natl.Acad. Sci. 20: 123 (1924). Type :— MEXICO, Sonora: C. V. Hartman 72 (Holotype at B destroyed. Lectotype (designated here): GH00071010 !) .

Quercus duraznillo f. pinetorum Trel., Mem. Natl. Acad. Sci. 20: 123, pl. 221b (1924). Type :— MEXICO, Sonora: C. V. Hartman 131 (Holotype: GH00055597 !) .

Quercus hastata Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 171 (1854) . Type:— U.S.A., Texas: C. Wright 666 (Holotype: K000832269 !; isotypes: G!, NY!, OXF!, US!). Liebmann (1854, p. 171) wrote “ Coll. Chr. Wright n. 666. (hb. Hook.!)” although the specimen marked as the holotype at K has a “Herbarium Benthamianum” stamp, it does have a determination in Liebmann’s hand.

Distribution: Mexico (Baja California, Chihuahua, Coahuila, Durango, Jalisco, Nuevo León, San Luis Potosí, Sonora,

Tamaulipas, Zacatecas), U.S.A. (Arizona, New Mexico, Texas) .

Quercus engelmannii Greene, Ill. W. Amer. Oaks 1: [33], pl. 15 figs. 2 & 3, pl. 17 (1889). Type:— U.S.A., California: E.L. Greene s. n. (16 Feb 1877) (Lectotype (designated here): NDG01270 !). It is uncertain whether there was ever an Albert Kellogg holotype of this species, but if there ever was one at CAS, it was not saved from the 1906 fire. The illustrations in Greene’s publication are by Kellogg. Greene collected this species in 1877 which is the one we are designating as the lectotype. Other collections by Greene at NDG were collected after the publication of the species, so the collection available to Greene at the time of the publication is the one appropriate for lectotypification.

Distribution: Mexico (Baja California), U.S.A. (California).

Quercus epileuca Trel., Mem. Natl. Acad. Sci. 20: 126, pl. 229a (1924). Type:— MEXICO, Sonora: C. V. Hartman 337 (Holotype: NY00253687 !; isotypes: A!, GH!, K!, UCSB (fragment from NY holotype)!, US!) .

Quercus hypoleuca Engelm. non Miq. (1858), Trans. Acad. Sci. St. Louis 3: 384 (1876). See Q. hypoleucoides below.

Quercus hypoleucoides A.Camus, Bull. Mus. Natl. Hist. Nat. , sér. 2, 4: 124 (1932). Type:— U.S.A., Arizona: A. Schott s. n. (28 Jul 1855) (Lectotype: NY00248698! designated by Trelease (1924, p. 223, pl. 226a) as the type of Q. hypoleuca Engelm. (1876) non Miq. (1855); isolectotypes: GH!, MO!). Nom. nov. for Q. hypoleuca Engelm (1876) non Miq. (1858). The type is similar to Q. scytophylla Liebm. , a species described from Oaxaca. Additional study is necessary to confirm our proposed synonymy.

Distribution: Mexico (Chihuahua, Coahuila, Durango, México, Sinaloa, Sonora, Zacatecas), U.S.A. (Arizona, New

Mexico, Texas).

Quercus flagellifera Trel., Mem. Natl. Acad. Sci. 20: 162, pl. 319 (1924). Type:— GUATEMALA, Alta Verapaz: O. F. Cook & R. F. Griggs 607 (Holotype: US 00089512!). This species is similar to Q. cortesii , and additional study may show that they should be treated as the same species.

Distribution: Guatemala.

Quercus frutex Trel., Mem. Natl.Acad. Sci. 20: 82, pl. 120a (1924). Type:— MEXICO, Ciudad de México: E. Bourgeau 68 (Holotype: C 10012503!; isotype: UCSB (fragment from C holotype)!).

Quercus frutex var. uhdeana Trel., Mem. Natl. Acad. Sci. 20: 83 (1924). Type:— MEXICO: Uhde 295 (Holotype: at B destroyed). Because no images or original material of Q. frutex var. uhdeana exist, the determination of this variety as probably Q. frutex is based on the Trelease (1924, p. 83) description and the fact that he also cited Uhde 295 as possibly a paratype of Q. vallicola (Trelease 1924, p. 81) which is a later homonym and treated here as a synonym of Q. frutex . It is also possible that Q. frutex var. uhdeana may be a synonym of Q. microphylla according to an anonymous reviewer (per. commun.).

Quercus intricata Trel. f. angusta Trel., Mem. Natl. Acad. Sci. 20: 85, pl. 127b (1924). Type:— MEXICO, Hidalgo: C. G. Pringle 13286 (Holotype: US00089548!; isotypes: CAS!, F!, GH!, LL (image)!, MEXU!, MICH!, TEX (image)!, UC!, VT!).

Quercus vallicola Trel. non Gand. (1890), Mem. Natl. Acad. Sci. 20: 81, pl. 116b (1924). Type:— Mexico, Ciudad de México: Uhde 296 (Holotype: B destroyed). There are apparently no isotypes, but because Q. vallicola Trel. is a later homonym, we are not lectotypifying the image of the holotype in Trelease (1924, pl. 116b). The placement of this species needs additional study, but we consider the type to fall within the variation found within Q. frutex as broadly interpreted.

Distribution: Mexico (Chihuahua, Ciudad de México, Guanajuato, Hidalgo, Jalisco, México, Michoacán, Oaxaca,

Puebla, Querétaro, San Luis Potosí, Sonora, Tlaxcala, Veracruz, Zacatecas).

Quercus fulva Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 183 (1854) . Type:— MEXICO, Sinaloa: B. Seemann 1973 (Holotype: K000512901 !; isotypes: BM!, C (as s.n.)!, GH!, UCSB (fragment from K holotype)!).

Quercus rosei Trel., Mem. Natl. Acad. Sci. 20: 137, pl. 260a (1924). Type:— MEXICO, Nayarit: J. N. Rose 3458 (Holotype: US 00089618!) .

Distribution: Mexico (Aguascalientes, Chihuahua, Coahuila, Durango, Guanajuato, Jalisco, Michoacán, Nayarit,

Nuevo León, Oaxaca, Querétaro, San Luis Potosí, Sinaloa, Sonora, Zacatecas).The occurrence of Q. fulva in Michoacán and Querétaro reported by Villaseñor (2016) needs to be confirmed.

Figures

FIGURE 1. Map showing the number of recognized Latin American Quercus species by country and for Mexico by state.