Xistera coventryi, Raven & Hebron & Williams, 2023

Raven, Robert J., Hebron, Wendy & Williams, Kylie, 2023, Revisions of Australian ground-hunting spiders VI: five new stripe-less miturgid genera and 48 new species (Miturgidae: Miturginae), Zootaxa 5358 (1), pp. 1-117 : 109-112

publication ID

https://doi.org/ 10.11646/zootaxa.5358.1.1

publication LSID

lsid:zoobank.org:pub:1A17A242-2E91-4F43-9E5D-063F8C0CBE72

DOI

https://doi.org/10.5281/zenodo.10249851

persistent identifier

https://treatment.plazi.org/id/7A20879E-5765-FF93-7DD4-FAE33B6A7217

treatment provided by

Plazi

scientific name

Xistera coventryi
status

sp. nov.

Xistera coventryi sp. nov.

Figs 2b, e, f View FIGURE 2 , 3a–c View FIGURE 3 , 85–86 View FIGURE 85 View FIGURE 86

Miturgidae QLD ”: Ramírez 2014.

Diagnosis. The darkest and amongst the smallest species of the genus, males differ from those of all other species in the RTA forming a short, rudimentary scoop ( Fig. 86b View FIGURE 86 ). Epigyne with two large, rounded lateral lobes.

Etymology. The species epithet is a patronym in honour of one of the collectors, A. John Coventry (1936– 2007), then Senior Curator of Herpetology at the National Museum of Victoria (now Museums Victoria), a highly respected scientist.

Type Material. Victoria: male holotype, 7.5 km ENE Kingower , 36°36’S 143°50 ‘E 26–29 Mar. 1990, A.J. Coventry, MV K4322 GoogleMaps .

Paratypes. Victoria: 1 ma., 4.6 km E of Hattah , 34°45’S 142°19’E, May 1986, A.L. Yen, MV K4487 GoogleMaps ; 4 ma., Mallee site K3, 7.5 km ENE of Kingower, 36°36’S 143°50’E, 26–29 Mar 1990, J. Coventry, MV K4322 GoogleMaps ; 4 ma., 3.9 km NE of Hattah , 34°45’S 142°19’E, May 1986, A.L. Yen, MV K4778 GoogleMaps ; 2 ma., Mallee site K1, 7 km ENE of Kingower, 36°34’S 143°50’E, 26–29 Mar 1990, J. Coventry, MV K4323 GoogleMaps ; 1 ma., 22.7 km N of Millewa Sth Bore , 34°34’S 141°02’E, Jun 1986, A.L. Yen, MV K4708 GoogleMaps ; 1 ma., Mallee Site K14, 7 km SSE of Kingower, 36°36’S 143°50’E, 26–29 Mar 1990, A.J. Coventry, MV K4307 GoogleMaps .

New South Wales: Gubatta, pitfall, 24–28 Feb. 1999, D. Driscoll: 1 ma., (site 2G), 33°38’S, 146°33’E, QM S51556 GoogleMaps ; 1 ma., (site 3G), 33°35’S, 146°35’E, QM S52344 GoogleMaps . Pulletop, pitfall, 24–28 Feb 1999, D. Driscoll: 2 ma., (site 1P), 33°58’S, 146°06’E, QM S51576 GoogleMaps ; 1 ma., (site 3P), 34°01’S, 146°04’E, QM S51569 GoogleMaps ; 1 ma., (site 3P), 34°01’S, 146°04’E, QM S51536 GoogleMaps ; 3 ma., (site 5P), 34°00’S, 146°05’E, QM S50987 GoogleMaps . Taleeban, 33°58’S, 146°26’E, pitfall, 23–27 Feb. 1999, D. Driscoll: 2 ma., (site 3 T), QM S50922 , S52345 GoogleMaps ; 1 ma., (site 4 T), 33°57’S, 146°27’E, QM S52394 GoogleMaps . 1 fem., Pulletop (site 8P), 34°01’S, 146°40’E, spinifex, pitfall, 12–18 Oct 1999, D. Driscoll, QM S52942 GoogleMaps .

Queensland: “Merigol”, 26°48’S, 145°49’E, mulga, pitfall, Apr. 2001, T GoogleMaps . Beutel : 1 ma., site 10, trap 15, QM S63392 ; 1 ma., site 12, trap 10, QM S63387 ; 1 ma., site 12, trap 5, QM S70025 ; 1 ma., site 12, trap 6, QM S70027 ; 1 ma., site 2, trap 7, 26°47’S, 145°49’E, mulga, pitfall, Apr. 2001, T GoogleMaps . Beutel , QM S63393 ; 1 ma., same data but site 3, trap 13, 26°47’S, 145°49’E, QM S63389 GoogleMaps ; 1 ma., same data but site 3, trap 14, QM S63396 GoogleMaps ; 1 ma., same data but site 4, trap 2, QM S63386 GoogleMaps ; 1 ma., site 7, trap 4, QM S63388 ; 1 ma., site 8, trap 4, QM S70029 ; 1 ma., site 8, trap 5, QM S70026 ; 2 ma., same data but site 6, trap 14, QM S70028 ; 1 ma., site 7, trap 15, QM S63383 ; 1 ma., site 9, trap 10, 26°49’S, 145°49’E, QM S63390 GoogleMaps ; 1 ma., same data but site 9, trap 7, QM S63395 GoogleMaps . 3 ma., Osbourne Mine site, SSE Mt Isa, Site 4A, 22°07’S, 140°34’E, pitfall, received Dec 1996, QM S53611 GoogleMaps ; 1 ma., Fleurs Stn (tree clearing), Summer 1998, T . Churchill , QM S106969 ; 2 ma., White Mts NP, 20°27’01”S 114°48’50”E, Royal Geographical Society , 18 Apr 2000, QM S72242 GoogleMaps ; 2 ma., Cooloola Village turnoff ( CV1 ), 26°05’S, 153°05’E, CSIRO GoogleMaps , Pitfall Trap, Aug-74, QM S53396 1 ma., “ Mt Tenison-Woods , 27°18’S, 152°45’E, 760 m, Pitfall Trap, 22 Jan–8 Apr 1975, QM S39594 GoogleMaps .

South Australia: 5 ma., Calperum Homestead , 21k W by N, 33°59’S, 140°30’E, mallee, pitfall & intercept traps, 1–30 Mar. 1995, K. R. Pullen , QM S41777 GoogleMaps ; 5 ma., Renmark , 14k WNW, 34°07’S, 140°37’E, mallee on dune, pitfall, 2 May–7 Jun. 1995, K. R. Pullen , QM S39081 , S116677 GoogleMaps ; 9 ma., Renmark , 31 km NW, 33°59’S, 140°30’E, mallee, flight/ ground intercept trap, 30 Mar.–2 May 1995, K. R. Pullen , QM S35779 GoogleMaps ; 10 ma., SDB 32 km N, 33°53’S, 140°44’E, chenopod scrubland, 29 Mar–3 May 1995, QM S39098 , QM S39097 GoogleMaps ; 17 ma., SDB 79 k NNW, 33°31’S, 140°24’E, Casuarina woodland, QM S35778 GoogleMaps ; 2 ma., SDB 3 May–6 Jun 1995, A. Lambie, QM S41779 ; 3 ma., Weetootla well, 30°29’S 139°14’E, NN1998.386-88; GoogleMaps 1 ma., Murray Mallee, Brookfield CP, 34°22’S, 139°30’E, early 1990, T.A. Morley , SAM NN1998.53 View Materials GoogleMaps ; 2 ma., Canopus, ca. 65 km N. of Renmark, Danggali Conservation Park , 33°31’S, 140°42’E, 7–18 Apr 1986, S.A. Collins (via A.D. Austin), Pan Trap, WAM98 View Materials /809–811. GoogleMaps

Description. Male, holotype MV K4322

Carapace 2.04 long, 1.58 wide. Opisthosoma 2.04 long, 1.13 wide. Total, 4.08.

I: 2.90, 1.13, 2.67, 2.47, 1.30, 10.47. II: 2.80, 1.27, 2.47, 2.20, 1.67, 10.41. III: 2.74, 1.07, 2.67, 2.86, 2.07, 11.41. IV: 3.54, 1.13, 3.34, 4.40, 1.80, 14.21. Palp: 0.87, 0.50, 0.50, -, 1.57, 3.44.

Colour. Dark spiders; carapace yellow brown centrally surrounded by wide dark margins and black edging around strial edges distally, along back caput margin and up to PLE. Opisthosoma dorsally gray with two pallid spots on anterior shoulders, some light flecking down and two large pallid spots distally; venter pallid with dark marks near spinnerets. Femora mottled dark brown and yellow with dense pile of light hairs and short dark bristles.

Carapace: With white hairs centrally from PME to just posterior to caput; black setae and some silver hairs in wide marginal zone. Fovea from caput base over carapace slope.

Eyes. Eight in 3 recurved rows; from above almost in 4 rows—front edge of ALE cuts base of back edge of AME, likewise in back row. All eyes of similar size but ALE smallest. PLE behind ALE and look sideways, back and up. Group front width, back width, length, 31:41:30. Sizes: AME, 8; ALE, 6; PME, 9; PLE, 9. AME–AME, 5, AME–ALE, 1; PME–PME, 3; PME–PLE, 7.

Chelicerae. Both margins with 2 teeth. sternum subcircular with pallid setae directed centrally from all corners. Palpal coxae. Anterior half pallid with shallow groove behind it. Labium. With groove around anterior edge.

Legs. with dense pile of hair. Trochanter notches ca. x 2, wider than deep. STC with 4–6 long teeth. Scopula: thin bipartite for 2/3–3/4 of metatarsi I, II, distal clusters on III, IV; denser but still thin on tarsi I–IV, extend just beyond tip as pseudotufts. Spines. I: fe pv1p1d3r2; pa 0; ti p2r2v2.2.2 me p1. r1. v2. II: fe pv1p1d3r2; pa 0; ti p2r2v2.2.2; me p1. r1v2.1. III: fe p3d3r3; pa 0; ti p2d2. r2v2.2.2; me p1.2.1r1.2.1v2.2.1. IV: fe p4d3r2; pa 0; ti p2d2.1r2v2.2.2; me p1.1.2r2.1.1.1. v2.2.1. Palp: fe d1.1; pa d1.1 (including 1 apical); ti p5d2; cymbium basodorsal 2.

Spinnerets. Both PLS and ALS long cylinders; PLS about 1/2 diameter of ALS. Apical PLS short, domed; PMS, shorter cylinders.

Palp. Tibia short with proventral keel and apical wedge and glabrous depression ventrally to retroedge. RTA saddle-like; from side basally two short rounded processes from chelae distally a groove of pallid unsclerotised tissue; saddle reflexes ventrally in wide arc and apical is rounded process with lateral fold. Cymbium short; margins narrow, retrobasal edge concealed by tegulum; apical cone very short partially glabrous ventrally; retrolateral margin with slight discontinuity. Median apophysis a short heavily sclerotised blade-like triangle with apical beak-like hook; separated from embolus by narrow groove. Embolus origin probasal, folds directly back under edge of subtegulum terminating just behind median apophysis and cradled by narrow transverse unsclerotised folded conductor. Tegulum large retrolateral dome, extending over edge of cymbium retrobasally. Subtegulum small, narrow with basal and prodistal parts. Cymbium with light scopula in proventral corner in oval area for distal half; no other modified setae evident; 1 spine probasally and 1 retrodorsally on cymbium and cluster of thicker setae apically.

Female QM S52942

Carapace 3.20 long, 3.53 wide. Opisthosoma 3.53 long, 2.53 wide. Total length, 6.7.

I: 2.53; 1.13; 1.93; 1.80; 1.40; 8.80. II: 2.27; 1.13; 1.87; 1.67; 1.33; 8.27. III: 2.40; 1.13; 1.87; 2.07; 1.33; 8.80. IV: 3.20; 1.33; 2.73; 3.33; 1.33; 11.93. Palp: 1.80; 0.73; 0.53; --; 0.80; 3.87.

Colour. Carapace brownish yellow with dark edges and dark broad marginal band with lighter scalloping in it. Opisthosoma dorsally dark brown with no evident pattern; venter pallid. Legs brownish yellow with irregular dark areas on femora (N.B.: not annulations); patellae distoventrally dark; tibiae dorsally dark; metatarsi and tarsi fawn; palpal femora with irregular dark areas subdistally. Eyes. Of similar sizes; grate-shaped tapetum in PME; rows of PER separated by 0.5 diameter of PME. Spines. As for male but palp: fe d1.1.2; pa p1; ti p6; ta p1v2 subdistal. Spinnerets. ALS and PLS of similar size.

Epigyne (as in Fig. 85c, d View FIGURE 85 ). Externally with two large rounded lateral lobes; internally with large wide duct folding from copulatory opening and then back to fertilisation ducts.

Distribution. Known from South Australia, Victoria, New South Wales and Queensland with one doubtful record from just east of Brisbane.

MV

University of Montana Museum

QM

Queensland Museum

T

Tavera, Department of Geology and Geophysics

CSIRO

Australian National Fish Collection

R

Departamento de Geologia, Universidad de Chile

SAM

South African Museum

Kingdom

Animalia

Phylum

Arthropoda

Class

Arachnida

Order

Araneae

Family

Xenoctenidae

Genus

Xistera

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