Dichotomius (Dichotomius) quinquedens (Felsche, 1910)

Chamorro, William, Marin-Armijos, Diego, senjo, Angelico & Vaz-De-Mello, Fernando Z., 2019, Scarabaeinae dung beetles from Ecuador: a catalog, nomenclatural acts, and distribution records, ZooKeys 826, pp. 1-343 : 118-119

Dichotomius (Dichotomius) quinquedens (Felsche, 1910) Plate 27A

Pinotus quinquedens Felsche, 1910: 343 (original description. Type locality: Ecuador, Los Llanos).

Pinotus quinquedens : Gillet 1911a: 62 (complete list of species); Luederwaldt 1929: 35 (characters in key); Blackwelder 1944: 208 (list of species from Latin America).

Dichotomius quinquedens : Medina et al. 2001: 138 (cited for Colombia); Carvajal et al. 2011: 320-321 (cited for Ecuador); Krajcik 2012: 92 (complete list of species); Sarmiento-Garcés and Amat-García 2014: 53 (characters in key), 64 (diagnosis).

Dichotomius (Dichotomius) quinquedens : Chamorro et al. 2018: 95 (cited for Ecuador).

Types specimens.

Pinotus quinquedens Felsche, 1910. Seven syntypes examined deposited at the SMTD. Lectotype to be designated in a future work on this species group.

Syntype (♂): "Los Lanos / Ecuador [p, black margin]", "quinquendens- / tatus Felsche / Ecuador [hw, purpure margin]", "Typus. [p, red label]", "Coll. C. Felsche / Kauf 20, 1918 [p, green label, black margin]".

Syntype (♂): "Los Lanos / Ecuador [p, black margin]", "Coll. C. Felsche / Kauf 20, 1918 [p, green label, black margin]".

Syntype (♀): "Los Lanos / Ecuador [p, red label, black margin]", "Coll. C. Felsche / Kauf 20, 1918 [p, green label, black margin]".

Syntype (♂): "Los Lanos / Ecuador [p, black margin]", "Coll. C. Felsche / Kauf 20, 1918 [p, green label, black margin]".

Syntype (♂): "Los Lanos / Ecuador [p, black margin]", "Coll. C. Felsche / Kauf 20, 1918 [p, green label, black margin]".

Syntype (♂): "Los Lanos / Ecuador [p, black margin]", "Coll. C. Felsche / Kauf 20, 1918 [p, green label, black margin]".

Syntype (♂): "Los Lanos / Ecuador [p, black margin]", "Coll. C. Felsche / Kauf 20, 1918 [p, green label, black margin]".

Distribution.

Colombia and Ecuador.

Records examined.

BOLIVAR: Bosque Protector Filo Palanga, 970 m (1 specimen MUTPL). CAÑAR: Cochancay, 1000 m (4 specimens CEMT); Javín, 850-1400 m (4 specimens CEMT); Joyapal (1 specimen MQCAZ). COTOPAXI: 4 km de Guasaganda, 500 m (2 specimens MQCAZ); Bosque Integral Otonga, 1800 m (2 specimens CEMT; 7 specimens MQCAZ); Los Libres, 2015 m (1 specimen MQCAZ); San Francisco de Las Pampa, 1500 m (7 specimens CEMT). EL ORO: Salvias Río Elvira, 1180 m (2 specimens MUTPL); Reserva Jocotoco, 1250 m (1 specimen MQCAZ). ESMERALDAS: El Placer, road Ibarra-San Lorenzo, 670 m (2 specimens MQCAZ); Estación Biológica Bilsa, 500 m (1 specimen MEPN); Playa de Oro, Estero Pote, 200 m (1 specimen CEMT). IMBABURA: Lita (2 specimens CEMT); Santa Cecilia (1 specimen QCAZ). MANABÍ: Puerto López, Guale, 200 m (1 specimen MUTPL); Puerto López, San Sebastian, 480 m (1 specimen MUTPL). PICHINCHA: Choconde San Miguel de los Bancos, 1200 m (4 specimens MUTPL); Chespi, Bellavista, 1380 (4 specimens MUTPL); Chiriboga, 1500 m (1 specimen MQCAZ); Estación Biológica Maquipucuna, 1650 (2 specimens MUTPL); Estación Biológica la Hesperia, 1200 m (2 specimens CEMT); Mindo, 1500 m (1 specimen MQCAZ); Palmeras (1 specimen MQCAZ); Pampas Argentinas (1 specimen MQCAZ). SANTO DOMINGO DE LOS TSÁCHILAS: Río Toachi (1 specimen MQCAZ); Santo Domingo (2 specimens CEMT). UNDETERMINED PROVINCE: Los Lanos [= Los Llanos] (7 specimens SMTD).

Temporal data.

Collected in all months except September.

Remarks.

Inhabits coastal lowland evergreen forests and coastal evergreen foothill forests from 200-1300 m a.s.l. In the Andean region, it was registered for the evergreen lower montane forests from 1500-1800 m a.s.l. Collected manually and with pitfall traps baited with human feces.