Dryinus Latreille, 1804

Olmi, Massimo, Guglielmino, Adalgisa & Vollaro, Massimo, 2011, Revision of fossil species of Dryinus belonging to constans group, with description of a new species (Hymenoptera: Dryinidae), Zootaxa 2981, pp. 43-55 : 44

Genus Dryinus Latreille, 1804

Dryinus Latreille 1804: 176 . Type species: Dryinus collaris ( Linnaeus 1767) , by first included species ( Latreille 1805).

Campylonyx Westwood 1835: 52 (synonymized by Olmi 1984); type species: Campylonyx ampuliciformis Westwood 1835, by monotypy.

Chelothelius Reinhard 1863: 409 (synonymized by Olmi 1993); type species: Chelothelius gryps Reinhard 1863, by monotypy.

Paradryinus Perkins 1905: 53 (synonymized by Olmi 1984); type species: Paradryinus koebelei Perkins 1905 , by original designation.

Chlorodryinus Perkins 1905: 57 (synonymized by Olmi 1984); type species: Chlorodryinus pallidus Perkins 1905 , by original designation.

Plastodryinus Kieffer in Kieffer & Marshall 1906: 496 (synonymized by Móczár 1965); type species: Plastodryinus szepligetii (Kieffer 1906), by original designation.

Mesodryinus Kieffer in Kieffer & Marshall 1906: 497 (synonymized by Olmi 1993); type species: Mesodryinus niger ( Kieffer 1904) , by original designation.

Hesperodryinus Perkins 1907: 40 (synonymized by Olmi 1984); type species: Hesperodryinus arizonicus Perkins 1907 , by original designation.

Perodryinus Perkins 1907: 43 (synonymized by Olmi 1993); type species: Perodryinus amoenus Perkins 1907 , by monotypy.

Lestodryinus Kieffer 1911: 108 (new name for Dryinus Latreille ; synonymized by Richards 1937); type species: Lestodryinus collaris ( Linnaeus 1767) , by automatic designation (see Dryinus Latreille 1804 ).

Tridryinus Kieffer 1913: 325 (synonymized by Olmi 1993); type species: Tridryinus striaticeps ( Kieffer 1909), by original designation.

Bocchoides Benoit 1954: 424 (synonymized by Olmi 1993); type species: Bocchoides bekilyensis Benoit 1954, by monotypy.

Richardsidryinus Móczár 1965: 376 (synonymized by Olmi 1991); type species: Richardsidryinus corsicus ( Marshall 1874), by original designation.

Avodryinus Ponomarenko 1981: 143 (synonymized by Olmi 1984); type species: Avodryinus canadensis Ponomarenko 1981, by monotypy and original designation.

Alphadryinus Olmi 1984: 973 (synonymized by Olmi 1993); type species: Alphadryinus asiaticus Olmi 1984 , by original designation.

Diagnosis. Female: macropterous; palpal formula 6/3; mandible with 1–4 teeth; occipital carina complete, or incomplete, or absent; antenna without tufts of long hairs on segments 5–10, usually with rhinaria, occasionally without; antennal segment 3 less than five times as long as segment 2; occasionally antennal segment 3 more than five times as long as segment 2 (in this case, notauli occasionally complete and scutum completely sculptured by numerous and parallel longitudinal keels); pronotal tubercle reaching or not tegula; fore wing with three cells enclosed by pigmented veins (costal, median and submedian); protarsus chelate; chela with rudimentary claw; segment 5 of protarsus less than twice as broad as enlarged claw; enlarged claw as long as, or shorter than protibia; tibial spurs 1/1/2, rarely 1/1/1. Male: macropterous; mandible with 1–3 teeth; palpal formula 6/3; occipital carina complete or incomplete; lateral regions of prothorax not continuous with mesopleura; epicnemium distinct; mesosternum fused with mesopleura and not distinct; fore wing with three cells enclosed by pigmented veins (costal, median and submedian); paramere without dorsal process; tibial spurs 1/1/2.

Distribution. Worldwide.

Hosts. Acanaloniidae , Cixiidae , Dictyopharidae , Flatidae , Fulgoridae , Issidae , Lophopidae , Ricaniidae , Tropiduchidae ( Guglielmino & Olmi 1997, 2006, 2007).

Species included. Two hundred and seventy-nine.

Remarks. The Neotropical species of Dryinus are divided into four groups ( Olmi 1993).