Agrostocynips Diaz

Buffington, Matthew L. & Scheffer, Sonja J., 2008, North American species of Agrostocynips Diaz (Hymenoptera: Figitidae: Eucoilinae), parasitoids of Agromyzidae (Diptera): bionomics and taxonomy, Zootaxa 1817, pp. 39-48 : 41-43

publication ID	https://doi.org/ 10.5281/zenodo.274371
DOI	https://doi.org/10.5281/zenodo.6228370
persistent identifier	https://treatment.plazi.org/id/7B189554-5239-DA07-FF44-F90E9B64FE2A
treatment provided by	Plazi
scientific name	Agrostocynips Diaz
status

Treatment

Agrostocynips Diaz

Agrostocynips Diaz, 1976: 32 . Type-species Agrostocynips clavatus Diaz , by original designation.

Diagnosis. Genal carina reduced. Orbital furrows reduced to absent. Mesoscutal keel absent. Scutellar plate with distinct tubercles present. Most easily confused with Zaeucoila and Aegeseucoela , both of which have complete genal carinae, a mesoscutal keel present (at least anteriorly) and distinct orbital furrows.

Redescription. Head. Nearly glabrous with scattered setae along lower face, clypeus and gena; ocellar hair patch absent ( Fig. 1 C View FIGURE 1. A – F ). Ventral 1/4 of lower face with admedian clypeal furrows converging toward the clypeus. Orbital furrows faint to completely reduced, originating from the lateral aspect of torulus and running to dorsal margin of malar sulcus ( Fig. 1 C View FIGURE 1. A – F ). Malar sulcus compound or simple. Malar space smooth; anteroventral margin with a raised protuberance. Genal carina present only along ventral margin of malar space ( Fig. 1 D View FIGURE 1. A – F ) (often visible only when head is removed from mesosoma).

Antennae. Female: 13 segments, moniliform ( Figs 1 A View FIGURE 1. A – F and 2 A View FIGURE 2. A – D ); segments 3-13 of sub-equal size; rhinaria present on segments 4-13. Male: 15 segments, moniliform; rhinaria present on segments 3-15; segments 4-15 sub-equal in size; segment 3 modified, slightly longer than segment 4, curved outwardly, excavated laterally.

Pronotum. Pronotal plate wide, with setae along the dorsal margin ( Fig. 1 F View FIGURE 1. A – F ); dorsal margin rounded; pronotal fovea open. Pronotal triangle absent. Pronotal impression absent. Lateral pronotal carina absent. Lateral portion of the pronotum smooth and glabrous ( Fig 1 E View FIGURE 1. A – F ).

Mesoscutum. Smooth and glabrous; no sculpture present ( Fig. 1 F View FIGURE 1. A – F ). Parascutal impression incomplete, narrow. Notauli, mesoscutal keel, parapsidal ridges and parapsidal hair line absent ( Fig. 1 F View FIGURE 1. A – F ).

Mesopectus. Upper part and lower part of mesopleuron smooth and glabrous ( Fig. 1 E View FIGURE 1. A – F ). Dorsal margin of mesopleural triangle well defined, rounded ventrally. Mesopleural carina simple. Lower part of mesopleuron bounded by distinct precoxal carina; surcoxal depression present, smooth.

Scutellum. Scutellar plate large to medium; midpit placed between center point of plate and posterior margin of plate; rim of plate translucent; prominent tubercles commonly found along the entire rim (often resembling "sawblade teeth" in lateral view) ( Figs 1 B, E–F View FIGURE 1. A – F , 2B View FIGURE 2. A – D ). Dorsal surface of scutellum reticulate, margined laterally and posteriorly; rounded laterally and posteriorly; laterodorsal and posterior projections of the scutellum absent. Lateral bars as long as wide; ventral lobe present. Scutellar fovea oval, smooth and deep.

Metapectal-Propodeal Complex. Anterior 3/4 of metapectus glabrous, posterior 1/4 setose ( Fig. 1 E View FIGURE 1. A – F ). Spiracular groove with a well defined dorsal margin, reduced ventral margin. Posterior margin of metapectus ridged. Metapleural ridge reduced to absent; submetapleural ridge absent. Anterior impressions of metepimeron and metepisternum present. Anteroventral cavity oval, setose. Propodeum covered in both long and short setae. Lateral propodeal carinae semi-parallel, bowed at junction with the auxiliary propodeal carinae; auxiliary propodeal carinae reduced. Nucha glabrous, crenulate.

Wings. Hyaline, with base of wing rarely darkened; setose ( Fig. 2 D View FIGURE 2. A – D –E). R1 complete; marginal cell as long as deep. Apical fringe present, medium in length.

Legs. Fore- and mid-coxa sub-equal in size, hind-coxa twice the size of either fore- or mid-coxa ( Fig. 2 B View FIGURE 2. A – D ). Fore coxa variously setose; mid and hind coxa with distinct lateral and posterior dorsoventral setal bands. Femora and tibiae with sparse setal lines; tarsomeres with dense appressed setae. Length of hind tarsomere 1 equal to the combined length of remaining hind tarsal segments.

Metasoma. Female: sub-equal in size to mesosoma ( Figs. 1 A View FIGURE 1. A – F and 2 A View FIGURE 2. A – D ). Base of syntergum with hairy ring present, composed of dense, short setae and longer, thin setae; remainder of metasoma glabrous. Micropunctures present on posterior 1/3 to 1/4 of syntergum, and on remaining terga. Terga posterior to syntergum abruptly directed ventrally, resulting in a near 90 degree angle between syntergum and terga. Male: As in female.

Distribution. Neotropical Region: Argentina, Chile, though Central America and into Central Mexico; Nearctic Region: Continental United States and into Southern Canada (British Columbia).

Biology. Agrostocynips clavatus has been recorded in the neotropics from several agromyzid species in the genera Melanagromyza ( De Santis et al. 1976) and Liriomyza ( Diaz & Valladares, 1979; Salvo, pers. comm.). Nearctic species of Agrostocynips have been reared from agromyzids in Agromyza (on Panicum ( Poaceae )), Liriomyza (several host plants) and from Phytomyza (on Ilex cassine and I. myrtifolia ).

Included species

clavatus Diaz, 1976: 32 . Holotype in MLP (not seen).

diastrophi (Ashmead) , Buffington (2004). Ganaspis diastrophi Ashmead, 1896: 184 -185. Holotype in USNM.

enneatoma ( Diaz) , Diaz & Gallardo (1997). Zaeucoila enneatoma Diaz, 1975: 1999 . Holotype in MLP (not seen).

robusta (Ashmead) , Buffington (2004). Chrestosema robusta Ashmead, 1894: 68 . Holotype in USNM.

Kingdom	Animalia
Phylum	Arthropoda
Class	Insecta
Order	Hymenoptera
Family	Figitidae