Toxognathus beauchenei Fleutiaux, 1918

Rosa, Simone Policena, 2014, Phylogenetic analysis and taxonomic revision of Physodactylinae (Coleoptera, Elateridae), Papéis Avulsos de Zoologia (São Paulo) 54 (18), pp. 217-292 : 240-241

Toxognathus beauchenei Fleutiaux, 1918

( Figs. 7 E-K; 34G)

Toxognathus beauchenei Fleutiaux, 1918a: 276 ; Fleutiaux, 1924: 178; Schenkling, 1927: 509; Fleutiaux, 1940a: 35.

Redescription (male, Fig. 34G ): Integument dark reddish-brown, antennae evenly brown or with antennomeres I-V yellow, VI-X black and XI reddish-brown. Total length 9.0- 9.5 mm; elytral base 1.0-1.05x as wide as pronotum, elytra 2.60-2.66x times longer than pronotum. Frons deeply concave along the anterior margin; frontal carina complete. Antennae ( Fig. 7E ) reaching the posterior angles of pronotum. Mandible unidentate. Pronotum weakly convex 1.19-1.22x wider than long with sides sinuous converging abruptly anteriad from anterior half, lateral carina directed ventrad anteriorly, posterior angles divergent and convex; punctures coalescent forming weak longitudinal striae. Elytral interstices costate. Free margin of metacoxal plate absent, tibia slightly widened apicad ( Figs. 7 G-I), outer angle of protibia truncate. Tergite IX with anterior margin straight or bisinuous.

Aedeagus ( Figs. 7 J-K): Phallobase 0.42x total length of aedeagus, 1.3x longer than wide, length ratio between lateroposterior and median parts 5.4; paramere with ventral anteromedian margin narrow, elongate and convergent, basal strut 0.29x total length of penis; ventral sclerite linear.

Holotype: [Hanoi, Tonkin], [ Toxognathus Beauchnei, Fleut. Type, Colletion Fleutiaux], [Type], [Muséum Paris, Coll. E. Fleutiaux], [ Toxognathus beauchenei (Fleutiaux) , J. Chassain det. 05], [Lectotype] (MNHN).

Material examined: VIETNAM. Tonkin , 1 ex. ( MNHN); Tam-Dao, 1 ex. ( MNHN) .

Distribution: VIETNAM.

Remarks: The specimen from Tam-Dao is somewhat different from the holotype in its more convex body, black integument with antennae and legs reddish-brown and longer antennae (surpassing the posterior angles by one antennomere). Its phallobase differs from that of the specimen from Tonkin in the relative ratio between length and width (0.98) and ratio between lateroposterior and median parts (7.3). Toxognathus beauchenei is most similar to T. costulatus by its size, color of integument and the costate elytral interstices, but can be easily distinguished by its protibia with dorsal apical angle truncate and not produced.

Figures

FIGURE 7: Toxognathus bakeri. A, pronotum; B, metaventrite; C, D, aedeagus (dorsal, ventral). Toxognathus beauchenei. E, antenna; F,hind wing; G, H, I, pro-, meso- and metathoracic leg (without tarsi); J, K, aedeagus (dorsal, ventral).

FIGURE 34: Habitus. A, Margogastrius schneideri (13.0 mm); B, Teslasena femoralis (9.0 mm); C, T. foucarti (9.5 mm); D, Idiotropia henoni (4.5 mm); E, Oligostethius capensis (10 mm); F,Toxognathus bakeri (6.0 mm); G, T. beauchenei (9.0 mm); H, T. coomani (6.5 mm); I, T. costulatus (11.0 mm); J, T. doherty (8.0 mm); K, T. fairmairei (7.5 mm); L, T. mouhoti (5.0 mm); M, Dactylophysus hirtus sp. nov. (10.5 mm); N, D. capixabensis nom. nov. (9.0 mm); O, D. tibialis (12.0 mm); P,Heterocrepidius mendax (9.0 mm); Q, Physodactylus asper sp. nov. (9.5 mm); R, P. besckei (11.0 mm). A, D = lectotypes; E-G, I, J, L, M, P, Q = holotypes.