Terebra neglecta Michelotti, 1847

Harzhauser, Mathias & Landau, Bernard M., 2023, The auger snails (Gastropoda, Conoidea, Terebridae) of the Miocene Paratethys Sea, Zootaxa 5385 (1), pp. 1-70 : 47-49

Terebra neglecta Michelotti, 1847

Figs 20A–C

* Terebra neglecta mihi — Michelotti, 1847: 214, pl. 17, fig. 8.

[ Terebrum] Terebrum neglectum (Micht.) — Sacco 1891a: 28, pl 1, fig. 63.

[ Terebrum] T [erebrum]. Neglectum var. expertusa Sacc. — Sacco 1891a: 28, pl 1, fig. 64.

Terebra cingulata (For.) — Sacco 1904: 106, pl. 23, fig. 33 [non Terebra cingulifera Lamarck, 1822 ].

Terebra neglecta Micht. — Friedberg 1911; 5, pl. 1, fig. 4.

Terebra neglecta Micht. — Friedberg 1938: 157.

Terebra (Terebra) neglecta Michelotti, 1847 — Glibert 1952a: 382, pl. 14, fig. 7.

Terebra neglecta Micht. — Csepreghy-Meznerics 1954: 57, pl. 7, figs 23 [non fig. 33, = Terebra tuberculifera Manzoni, 1870 ]. Terebra acuminata Borson — Csepreghy-Meznerics 1956: 436, pl. 12, fig. 12 [non Terebra acuminata Borson, 1820 ].

Terebra (Myurellina) neglecta Michelotti, 1847 — Glibert 1960: 113.

Terebra (Terebra) neglecta (Michelotti) 1847 —Kojumdgieva in Kojumdgieva & Strachimirov 1960: 217, pl. 51, fig. 8. Terebra neglecta Micht. — Báldi 1960: 87.

Terebra neglecta Micht. — Florei 1961: 692, pl. 9, fig. 70.

Terebra (Terebra) neglecta Michelotti, 1847 — Tejkal et al. 1967: 209, pl. 12B, fig. 3

[non Terebra acuminata Borson, 1820 ].

Terebra (T.) neglecta (Micht.) — Stancu & Andreescu 1968: 465, pl. 6, fig. 72.

Terebra (Myurellina) neglecta Michelotti — Davoli, 1977 p. 165, pl. 4, fig. 1.

Terebra (Terebra) pseudopertusa badeniana ssp. N. — Švagrovský 1982: 405, pl. 6, figs 1–2.

? Terebra (Myurellina) neglecta Michelotti — Ruggieri & Davoli 1984: 75, pl. 14, fig. 16.

Terebra (Myurellina) neglecta Michelotti, 1847 — Davoli 1990: 105, pl. 10, figs 5, 6, 9.

Terebra (Myurella) acuminata Borson, 1820 —Harzhauser 2002: 116, pl. 10, fig. 9 [non Terebra acuminata Borson, 1820 ]. Strioterebrum basteroti (Nyst, 1843) — Popa et al. 2014: 18, pl. 5, fig. 11 [non Strioterebrum basterotii Nyst , 184]. Strioterebrum acuminatum ( Borson, 1820) — Landau et al. 2020: 94, pl. 83, fig. 1 [non Terebra acuminata Borson, 1820 ].

non Terebra neglecta Nyst — Strausz 1962: 30, pl. 4, figs 36–37 [= Terebra acuminata Borson, 1820 ].

non Terebra neglecta Michelotti — Răileanu & Negulescu 1964: 188, pl. 15, fig. 6 [? Strioterebrum borianum ( Švagrovský, 1982) ].

non Terebra neglecta Micht. — Strausz 1954: 36, pl. 2, fig. 42 [= Terebra acuminata Borson, 1820 ].

non Terebra neglecta Michelotti, 1847 — Strausz 1966: 391, pl. 4, figs 36–37 [= Terebra acuminata Borson, 1820 ].

non Terebra neglecta (Micht.) — Csepreghy-Meznerics 1972: 34, pl. 18, fig. 8 [= Terebra tuberculifera Manzoni, 1870 ].

non Terebra neglecta (Michelotti) 1847 — Bohn-Havas 1973: 1072, pl. 8, fig. 8 [?= Terebra acuminata Borson, 1820 ].

non Terebra (Myurellina) acuminata neglecta Michelotti, 1847 —A.W. Janssen 1984: 340, pl. 13, fig. 11, pl. 77, figs 9–10 [= Terebra acuminata Borson, 1820 ].

non Terebra (Terebra) neglecta Michelotti — Schultz 1998: 72, pl. 29, fig. 15 [ Terebra tuberculifera Manzoni 1870 ].

Type material. Syntype illustrated in Michelotti (1847: pl. 17, fig. 8); Tortona , Italy, Tortonian (Late Miocene). We are not aware of the whereabouts of the material .

Illustrated material. NHMW 1861/0033/0020a, SL: 42.2 mm, MD: 8.6 mm, Bujtur ( Romania), Figs 20A 1 –A 2 . NHMW 1861/0033/0020b, SL: 46.5 mm, MD: 9.6 mm, Bujtur ( Romania), Figs 20B 1 –B 2 . NHMW 1861/0033/0020c, SL: 43.5 mm, MD: 9.1 mm, Bujtur ( Romania), Figs 20C 1 –C 2 .

Additional material. 40 spec., NHMW 2013 /0078/0410, Baden ( Austria) ; 3 spec., NHMW 2013 /0078/0412, Baden ( Austria) ; 25 spec., NHMW 1861 /0033/0020, Bujtur ( Romania) ; 6 spec., NHMW 1862 /0033/0013, Pöls ( Austria) ; 5 spec., NHMW 1868 /0001/0415, Lăpugiu de Sus ( Romania) ; 3 spec., NHMW 2016 /0177/0942, Lăpugiu de Sus ( Romania) ; 4 spec., NHMW 2013 /0300/0396, Baden Sooss ( Austria) ; 2 spec., NHMW 2013 /0078/0411, Baden ( Austria) ; 5 spec., NHMW 1997 z0178/1202, Bad Vöslau ( Austria) ; 15 spec., NHMW 2018 /0248/0354, CoŞteiu de Sus ( Romania) ; 7 spec., NHMW 2018 /0248/0355, CoŞteiu de Sus ( Romania) ; 10 spec., NHMW 1846 /0037/0125, Gainfarn ( Austria) .

Revised description. Medium sized, slender shell with slightly gradate spire, of about 16 teleoconch whorls; apical angle ~20°. Protoconch unknown (from Paratethyan material). Early teleoconch whorls conical to subcylindrical with prominent orthocline axial ribs, separated by narrower interspaces. Subsutural band beaded, delimited by distinct groove. Later teleoconch whorls relatively low, subcylindrical with prominent, rounded to subquadratic beads on subsutural band, and broad, slightly prosocline axial ribs. Beads on subsutural band slightly shifted relative to axial ribs, typically situated between tips of axial ribs. Axial sculpture weakening on last whorl. Last whorl low, ~26% of total height, subcylindrical with rounded basal angulation. Base strongly constricted. Aperture moderately narrow. Fasciole broad, flat, smooth, adapically delimited by narrow carina. Columella twisted, excavated in adapical half, with two moderately prominent columellar folds. Anal canal indistinct. Outer lip thin. Siphonal canal moderately long, narrow, twisted, slightly deflected to the left.

Paratethyan synonyms. Terebra (Terebra) pseudopertusa badeniana Švagrovský 1982 ; holotype, Institute for Geology and Paleontology, PFUK, Bratislava, illustrated in Švagrovský 1982: pl. 6, fig. 1, Borský Mikuláš ( Czech Republic).

Discussion. Terebra neglecta is a poorly understood species because of the numerous misidentifications in the literature. It was often confused with Terebra acuminata Borson, 1820 .

The specimen illustrated by Michelotti (1847) is a moderately slender shell with a gradate spire and prominent axial ribs with the adapical tips being squeezed in between the concavities between the subquadratic beads of the subsutural band. This specimen does not differ from the Paratethyan shells illustrated herein and described by Švagrovský (1982) as Terebra (Terebra) pseudopertusa badeniana . Terebra neglecta differs from Terebra acuminata ( Borson, 1820) and Terebra tuberculifera Manzoni, 1870 in its distinctly less slender outline and a shorter spire and the prominent, regular axial sculpture. Moreover, it lacks the concavity of the whorls of Terebra tuberculifera .

Molecular data documented that Cinguloterebra Oyama, 1961 [type species Terebra hedleyana Pilsbry, 1905 , = Terebra adamsii E.A. Smith, 1873 ] is a junior synonym of Terebra ( Fedosov et al. 2020: 367) . Therefore, Cinguloterebra neglecta Poppe, Tagaro & Terryn, 2009 , established for a living species from the Philippines, became a secondary homonym of Terebra neglecta Michelotti, 1847 . To solve this problem, Terryn (2023) introduced Terebra philpoppei Terryn, 2023 as replacement name for the extant species.

Paleoenvironment. The species occurs in large numbers in clays of the Baden Formation, suggesting middle to outer neritic water depths ( Kranner et al. 2021), but is also found in coastal marine settings (e.g., Weinsteig, Harzhauser 2002).

Distribution in Central Paratethys. Karpatian (Early Miocene): Korneuburg Basin: Karnabrunn, Weinsteig ( Austria) ( Hoernes &Auinger 1880; Harzhauser 2002); North Alpine Foreland Basin: Slup ( Czech Republic) ( Tejkal et al. 1967). Badenian (Middle Miocene): Polish-Ukrainian Fore-Carpathian Basin: Dryszczów ( Poland) ( Friedberg 1911); Vienna Basin: Borský Mikuláš ( Czech Republic) ( Švagrovský 1982); Baden, Baden Sooss, Bad Vöslau, Gainfarn ( Austria) ( Hoernes &Auinger 1880; hoc opus); Eisenstadt-Sopron Basin: Forchtenau ( Austria) ( Hoernes & Auinger 1880); Styrian Basin: Pöls ( Austria) ( Hoernes & Auinger 1880); Cserhát Mountains: Mátraverebély ( Hungary) ( Csepreghy-Meznerics 1954); Făget Basin: Bujtur, CoŞteiu de Sus, Lăpugiu des Sus ( Romania) ( Hoernes & Auinger 1880; Popa et al. 2014; hoc opus). Caransebeş Basin: DelineŞti, Rugi ( Romania) ( Stancu & Andreescu 1968); Dacian Basin: Târnene, Staropatica ( Bulgaria) (Kojumdgieva in Kojumdgieva & Strachimirov 1960).

Proto-Mediterranean Sea. Burdigalian (Early Miocene): Colli Torinesi ( Italy) ( Sacco 1891a). Langhian (Middle Miocene): Colli Torinesi: Sciolze ( Sacco 1891a); Tortonian (Late Miocene): Po Basin: Montegibbio, Stazzano, S. Agata, Sogliano, Tortona ( Italy) ( Sacco 1891a, Davoli 1977, 1990).

Northeastern Atlantic. Langhian: Loire Basin ( France) ( Glibert 1952a); Tortonian (Late Miocene): Loire Basin ( France) ( Landau et al. 2020).

Figures

FIGURE 1. Measurements for shell morphology, spire profiles and types of siphonal canals; AA: apical angle, MD: maxium diameter.

FIGURE 2. Fusoterebra terebrina (Bellardi & Michelotti, 1840). A1–A2. NHMW 1997z/0178/1787, Walbersdorf (Austria). B1–B2. NHMW 1997z/0178/1787, Walbersdorf (Austria). C1–C2. NHMW 1997z/0178/1787, Walbersdorf (Austria). D1–D2. NHMW 1846/0037/0121, Baden (Austria), lectotype of Terebra fusiformis Hörnes 1852.F. NHMW 1866/0001/1055, Forchtenau (Austria), holotype of Terebra bigranulata Hoernes & Auinger, 1880. E1–E2. NHMW 2023/0292/0001, Baden Sooss (Austria).

FIGURE 20. Terebra neglecta Michelotti, 1847. A1–A2. NHMW 1861/0033/0020a, Bujtur (Romania). B1–B2. NHMW 1861/0033/0020b, Bujtur (Romania). C1–C2. NHMW 1861/0033/0020c, Bujtur (Romania).