Fusoterebra terebrina ( Bellardi & Michelotti, 1840 )

Harzhauser, Mathias & Landau, Bernard M., 2023, The auger snails (Gastropoda, Conoidea, Terebridae) of the Miocene Paratethys Sea, Zootaxa 5385 (1), pp. 1-70 : 7-9

publication ID	https://doi.org/ 10.11646/zootaxa.5385.1.1
publication LSID	lsid:zoobank.org:pub:7FBEC9FB-E31E-4CA4-8BD0-BE6D35322C3E
DOI	https://doi.org/10.5281/zenodo.10459880
persistent identifier	https://treatment.plazi.org/id/7C37CE5F-FFA6-AB3D-FF2C-FE90302DF9E7
treatment provided by	Plazi
scientific name	Fusoterebra terebrina ( Bellardi & Michelotti, 1840 )
status

Treatment

Fusoterebra terebrina ( Bellardi & Michelotti, 1840)

Figs 2A–F View FIGURE 2 , 3A View FIGURE 3

* Fusus terebrinus Bon. — Bellardi & Michelotti 1840: 111, pl. 2, fig. 4.

Fusus terebrinus — Hörnes 1848: 19.

Terebra fusiformis Hörnes — Hörnes 1852: 135, pl. 11, figs 31a–b.

T [erebra]. bigranulata— Hoernes 1880: 246 [nomen nudum].

Terebra (Acus) fusiformis M. Hoern ,— Hoernes & Auinger 1880: 109.

Terebra (?) bigranulata View in CoL nov. form.— Hoernes & Auinger 1880: 111, pl. 12, figs 22a–c.

[ Terebra] Fusoterebra terebrina (Bon.)— Sacco 1891a: 60, pl. 2, fig. 70.

[Terebra] F [usoterebra]. terebrina var. pseudanodulosa Sacc. — Sacco 1891a: 60, pl. 2, fig. 71.

[Terebra] F [usoterebra]. terebrina var. unicarinata Sacc. — Sacco 1891a: 60, pl. 2, fig. 72.

[ Terebra] F [usoterebra]. terebrina var. diversenodulosa Sacc. — Sacco 1891a: 61, pl. 2, fig. 73.

[ Terebra] F [usoterebra]. terebrina var. ventresulcata Sacc. — Sacco 1891a: 61, pl. 2, fig. 74.

[ Terebra] F [usoterebra]. terebrina var. pernodulosa Sacc. — Sacco 1891a: 61, pl. 2, fig. 75.

Fusoterebra terebrina var. bigranulata (R. Hoern. et Auing.) — Sacco 1891a: 61.

Fusoterebra terebrina (Bon.) — Cossmann 1896: 57, pl. 4, fig. 14.

Strioterebrum (Fusoterebra) terebrinum (Bon.) — Wenz 1943: 1482, fig. 4177.

Terebra (Fusoterebra) terebrina Bon. — Korobkov 1955: plate captions, pl. 92, fig. 12.

S [trioterebrum]. (F [usoterebra].) terebrinum (Bon.) — Sieber 1958: 162.

S[trioterebrum]. (F [usoterebra].) terebrinum bigranulatum (R. Hörn, et Au.)— Sieber 1958: 162.

Terebra (Fusoterebra) terebrina Bonelli (in Bellardi & Michelotti), 1841) — Strausz 1966: 393, pl. 4, fig. 31, pl. 73, fig. 7. Strioterebrum (Fusoterebra) terebrinum (Bellardi & Michelotti) — Robba 1968: 612, pl. 46, figs 8–9.

Terebra (Fusoterebra) terebrina (Bon.) — Csepreghy-Meznerics 1969: 99, pl. 6, figs 14, 20.

Terebra (Fusoterebra) terebrina pernodulosa Sacco — Csepreghy-Meznerics 1969: 100, pl. 6, fig. 18.

Terebra (Fusoterebra) terebrina (Bon.) — Csepreghy-Meznerics 1972: 35, pl. 18, figs 1, 2.

Terebra (Fusoterebra) terebrina pernodulosa Sacco — Csepreghy-Meznerics 1972: 35, pl. 18, fig. 3.

Strioterebrum (Fusoterebra) terebrinum (Bellardi & Michelotti) — Davoli 1977: 155, pl. 2, figs 17–22.

Fusoterebra terebrina var. pseudanodulosa Sacco — Ferrero Mortara et al. 1984: 65, pl. 8, figs 6a–b.

Fusoterebra terebrina var. pernodulosa Sacco — Ferrero Mortara et al. 1984: 65, pl. 8, figs 11a–b.

Strioterebrum (Fusoterebra) terebrinum ( Bellardi & Michelotti, 1840) — Davoli 2003: 460, pl. 2, figs 9, 13, 15, pl. 3, fig. 10. Terebra (Fusoterebra) terebrina ( Bellardi & Michelotti, 1840, Bonelli m.s.)— Zunino & Pavia 2009: 365, pl. 2, fig. 9.

Type material. Syntype (?), BS.032.06.001, stored in the Museo Regionale di Scienze Naturali Torino ( Italy), illustrated in Bellardi & Michelotti (1840: pl. 2, fig. 4) and Sacco (1891a: pl. 2, fig. 71); Stazzano ( Italy), Tortonian (Late Miocene). The specimen is listed as syntype with question mark by Ferrero Mortara et al. (1984: 65).

Illustrated material. NHMW 1866/0001/1055, SL: 6.7 mm, MD: 2.6 mm, Forchtenau ( Austria), holotype of Terebra bigranulata Hoernes & Auinger, 1880 , Figs 2F View FIGURE 2 . NHMW 1846/0037/0121, SL: 29.4 mm, MD: 7.7 mm, Baden ( Austria), Lectotype (designated herein) of Terebra fusiformis Hörnes 1852 , illustrated in Hörnes (1852: pl. 11, fig. 31), Figs 2D View FIGURE 2 1 –D View FIGURE 1 2 View FIGURE 2 . NHMW 1997z/0178/1787, SL: 44.7 mm, MD: 11.0 mm, Walbersdorf ( Austria), Figs 2A View FIGURE 2 1 –A View FIGURE 1 2 View FIGURE 2 . NHMW 1997z/0178/1787, SL: 43.9 mm, MD: 10.5 mm, Walbersdorf ( Austria), Figs 2B View FIGURE 2 1 –B View FIGURE 1 2 View FIGURE 2 . NHMW 1997z/0178/1787, SL: 38.5 mm, MD: 9.1 mm, Walbersdorf ( Austria), Figs 2C View FIGURE 2 1 –C View FIGURE 1 2 View FIGURE 2 . NHMW 2023/0292/0001, SL: 40.8 mm, MD: 9.2 mm, Baden Sooss ( Austria), Figs 2E View FIGURE 2 1 –E View FIGURE 1 2 View FIGURE 2 . NHMW 2023/0293/0001, SL: 15.6 mm, MD: 3.7 mm, Baden ( Austria), Figs 3A View FIGURE 3 .

Additional material. 16 spec., NHMW 1846 View Materials /0037/0121, Baden and Möllersdorf ( Austria) , paratypes of Terebra fusiformis Hörnes 1852 ; 7 spec., NHMW 2023 View Materials /0294/0001, Walbersdorf ( Austria) ; 5 spec., NHMW 2013 View Materials /0300/0399, Baden-Sooss ( Austria) ; 1 spec., NHMW 1869 View Materials /0001/0067, Baden-Sooss ( Austria) ; 30 spec., NHMW 2013 View Materials /0300/0398, Baden-Sooss ( Austria) ; 14 spec., NHMW 2013 View Materials /0300/0418, Baden ( Austria) ; 3 spec., NHMW 1866 View Materials /0001/0175, Baden ( Austria) ; 8 spec., NHMW 2010 View Materials /0004/1097, Bad Vöslau ( Austria) ; 2 spec., NHMW 1863 View Materials /0015/0430 , 5 spec., NHMW 2012 View Materials /0229/0174, Traiskirchen ( Austria) ; 1 spec., NHMW 1862 View Materials /0029/0014, Rudice ( Czech Republic) ; 1 spec., NHMW 2016 View Materials /0177/0949, Lăpugiu de Sus ( Romania) ; 2 spec., NHMW 2016 View Materials /0177/0950, Lăpugiu de Sus ( Romania) ; 7 spec., NHMW 1858 View Materials /0043/0019, Lăpugiu de Sus ( Romania) , 2 spec., NHMW 1870 View Materials /0049/0004, CoŞteiu de Sus ( Romania) .

Revised description. Medium sized, slender fusiform shell of up to 13 teleoconch whorls; apical angle ~20– 22°. Protoconch broad conical of 2.5 convex whorls. First teleoconch whorl with mid-whorl angulation and weak subsutural band. Axial ribs form prominent beads on shoulder cord. Later whorls with beaded subsutural band, resulting in two beaded spiral cords. Indistinct axial ribs may form third row of weak beads just above abapical suture. Later teleoconch whorls flat sided. Sculpture of dense orthocline axial ribs separated by narrower interspaces. Axial ribs beaded on moderately prominent subsutural band, with second spiral row of beads on shoulder cord. Spire profile weakly gradate. Subsutural band and beads weaken on later whorls grading into weak axial ribs separated by wider interspaces.Shoulder cord forming periphery. On last whorl axial ribs weaken slightly, indistinct on the last half whorl in some specimens. Faint spiral threads in interspaces on some specimens. Last whorl high, ~37–43% of total height; slowly contracting into base. Fasciole weak with some more prominent growth increments. Aperture very narrow, elongate. Anal canal indistinct, weakly incised. Columella slightly twisted, excavated in adapical half. No columellar fold. Columellar callus forming broad rim, usually not distinctly delimited from base. Outer lip thin. Siphonal canal narrow, long, slightly twisted and deflected to the left.

Paratethyan synonyms. Specimens assigned herein to this species were described by Hörnes (1852) as Terebra fusiformis although he clearly stated that his specimens were identical to ‘ Fusus terebrinus Bellardi & Michelotti’. Hörnes (1852) even included specimens from Tortona in his description, but nevertheless introduced a new name for this species. Hörnes (1852) seems to have suppressed the older name because the curator Paul Maria Partsch (1791–1856) had labelled the Viennese specimens already in the early 1840ies as a new species in the NHMWcollection. However, Bellardi & Michelotti’s name clearly has priority.

Terebra bigranulata Hoernes & Auinger, 1880 is based on a juvenile specimen of Fusoterebra terebrina with prominent beads and at that stage of growth nearly absent axial ribs. Its close relation with Fusoterebra terebrina was already recognized by Sacco (1891a) and Davoli (1977).

Discussion. Fusoterebra terebrina is variable concerning its sculpture and especially the subsutural beads and the shoulder beads may be very prominent or almost subobsolete at variable stages of growth. In most Paratethyan specimens, delicate axial ribs are the predominant sculpture on late teleoconch whorls. Early whorls are reminiscent of some species of Duplicaria Dall, 1908 , such as Duplicaria herberti Malcolm, Terryn & Fedosov in Fedosov et al., 2020 and D. mozambiquensis Bratcher & Cernohorsky, 1982 , but the fusiform last whorl of F. terebrina excludes a closer relation (see Fedosov et al. 2020: figs 11A–D).

Paleoenvironment. The frequent occurrence in the middle Badenian Baden Formation in the Vienna Basin suggests middle to outer neritic water depths ( Kranner et al. 2021), which is in line with the preferred habitat of living pellifronine species.

Distribution in Central Paratethys. Badenian (Middle Miocene): North Alpine-Carpathian Foreland Basin: Rudice, Židlochovice ( Czech Republic) ( Hoernes & Auinger 1880); Vienna Basin: Baden, Möllersdorf ( Austria) ( Hoernes & Auinger 1880); Eisenstadt-Sopron Basin: Forchtenau ( Austria) ( Hoernes & Auinger 1880), Sopron ( Hungary) ( Strausz 1966); Bükk Mountains: Borsodbóta, Bóta ( Hungary) ( Csepreghy-Meznerics 1972); Făget Basin: CoŞteiu de Sus, Lăpugiu des Sus ( Romania) ( Hoernes & Auinger 1880).

Proto-Mediterranean Sea. Langhian (Middle Miocene): Colli Torinesi: Monte Cappuccini ( Italy) ( Zunino & Pavia 2009). Tortonian (Late Miocene): Po Basin: S. Agata, Giusulana, Montegibbio, Rio Mazzapiedi-Castellania, Stazzano ( Italy) ( Sacco 1891a; Robba 1968; Davoli 1977).

NHMW	Naturhistorisches Museum, Wien

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Taxonomy

Kingdom	Animalia
Phylum	Mollusca
Class	Gastropoda
Order	Neogastropoda
SuperFamily	Conoidea
Family	Terebridae
SubFamily	Pellifroniinae
Genus	Fusoterebra

1852

Fusus terebrinus — Hörnes 1848: 19 .

Hornes, M. 1848: 19

1848

Fusus terebrinus

Bellardi, L. & Michelotti, G. 1840: 111

1840

Fusoterebra terebrina ( Bellardi & Michelotti, 1840 )