Trachydonia Bernhauer, 1928

Hlaváč, Peter, Newton, Alfred F. & Maruyama, Munetoshi, 2011, World catalogue of the species of the tribe Lomechusini (Staphylinidae: Aleocharinae) 3075, Zootaxa 3075, pp. 1-151 : 81-84

Trachydonia Bernhauer

Zyras (Trachydonia) Bernhauer, 1928a: 42 . Type species: Zyras oxyteloides Bernhauer , by original designation Trachydonia Bernhauer : Kistner, 1997: 161 (separate genus); Kistner, 2003b (redescription)

actuosa Last, 1969a: 133 ( Zyras (Trachydonia), TL : Plato Bateke, Mbe) Distribution: Congo-Brazzaville aethiops Cameron, 1937b: 269 ( Zyras (Trachydonia), TL : Mayumbe, Buende-Suindi) Distribution: Dem. Rep.

Congo albomarginata Bernhauer, 1928a: 47 ( Zyras (Trachydonia), TL : Deutsch-Ostafrika, Lulanguru, W of Tabora)

Distribution: Tanzania, Ghana, Benin alboplagiata Bernhauer, 1928a: 49 ( Zyras (Trachydonia), TL : Deutsch-Ostafrika, Kwakiyembe) Distribution:

Congo-Brazzaville, Senegal, Tanzania, Togo, Ghana, Dem. Rep. Congo annosa Last, 1967: 110 ( Zyras (Trachydonia), TL : Angola, Mussangueje, 120 km SSW de Dundo) Distribution:

Angola, Ghana, Nigeria anommatis Last, 1977c: 941 ( Zyras (Trachydonia), TL : Zaïre, Stanleyville; HA: Dorylus (Anomma) nigricans wilverthi Emery ) Distribution: Dem. Rep. Congo arrogans Bernhauer, 1915c: 170 ( Zyras, TL : Deutsch-Ostafrika, Kilimandjaro) Distribution: Tanzania asperata Bernhauer, 1929c: 191 ( Zyras (Trachydonia), TL : Portugiesisch Guinea, Bolama) Distribution: Guinea

Bissau, Dem. Rep. Congo attestor Last, 1960c: 54 ( Zyras (Trachydonia), TL : Tanganyka Terr., Bunduki, Uluguru Mts., moyenne Mgeta,

1300 m) Distribution: Tanzania bayoni Bernhauer, 1929c: 189 ( Zyras (Trachydonia) ; TL: Uganda, Entebbe & Victoria Nyanza Arcip., di Sesse,

Bugala) Distribution: Uganda, Rwanda (?) bihamata Fauvel, 1907: 68 ( Zyras, TL : Mt. Nyro [ Kenya]; Ikutha [ Tanzania]; Abyssinie (Mansinam, Bogos, vers

1500m) [ Eritrea]; Zanzibar) Distribution: Eritrea, Kenya, Tanzania, Zanzibar bilimbata Pace, 1986b: 105 ( Zyras (Trachydonia) ; TL: Tanzania, Usa river, 3900 feet) Distribution: Tanzania cameroni Scheerpeltz, 1934: 1651 ( Zyras (Trachydonia) , new name for Zyras mimus Cameron, 1932 ) Distribution:

Dem. Rep. Congo

= mimus Cameron, 1932: 38 ( Zyras (Trachydonia), TL : Haut Uele, Moto; preoccupied in Zyras , not

Myrmedonia mima Péringuey, 1904 ) capiriensis Bernhauer, 1915h: 300 ( Zyras, TL : Kapiri) Distribution: Cameroon, Dem. Rep. Congo cicatricosa Bernhauer, 1915: 170 ( Zyras, TL : Abyssinien: Harrar) Distribution: Ethiopia, Sudan curticollis Bernhauer, 1928a: 46 ( Zyras (Trachydonia), TL : Deutsch-Ostafrika, Myombo & Narobi b. Tanga)

Distribution: Tanzania, Kenya densata Bernhauer, 1930a: 143 ( Zyras (Trachydonia), TL : Belgisch-Kongo, Haut Uele, Yebo Moto) Distribution:

Cameroon, Congo-Brazzaville, Dem. Rep. Congo, Angola domita Last, 1969a: 135 ( Zyras (Trachydonia), TL : Lafine reservation) Distribution: Congo-Brazzaville endroedyyoungai Last, 1973a: 154 ( Zyras (Trachydonia), TL : Ghana, Northern region, Banda-Nkwanta, 150 m)

Distribution: Ghana excellens Cameron, 1952a: 834 ( Zyras (Trachydonia), TL : Dahomey, Bassila) Distribution: Benin, Togo, Ghana,

Angola flavescens Cameron, 1939d: 22 ( Zyras (Trachydonia), TL : Djeroeklegi, Zuid Banjoemans) Distribution: Indonesia

(Java) frater Bernhauer, 1929c: 193 ( Zyras (Trachydonia), TL : Portugiesisch Guinea, Bolama) Distribution: Guinea

Bissau gaerdesi Scheerpeltz, 1966: 68 ( Zyras (Trachydonia), TL : Okahandja) Distribution: Namibia gelanor Tottenham, 1953 ( Zyras (Trachydonia), TL : Uganda: Kawanda, near Kampala) Distribution: Uganda gentilis Péringuey, 1904: 213 ( Myrmedonia, TL : Natal, Frere) Distribution: Rep. South Africa (Natal), new combination in Last, 1980b: 170 granulosa Bernhauer, 1943: 303 ( Zyras (Trachydonia), TL : Kesenyi) Distribution: Congo-Brazzaville, Dem. Rep.

Congo, Tanzania gridellii Bernhauer, 1928a: 48 ( Zyras (Trachydonia), TL : Portugiesisch Guinea, Bolama) Distribution: Guinea

Bissau hector Bernhauer, 1915c: 171 ( Zyras, TL : Deutsch-Ostafrika, Kaguru) Distribution: Tanzania hostis Last, 1969a: 134 ( Zyras (Trachydonia), TL : Brazzaville, Orstom park) Distribution: Congo-Brazzaville,

Tanzania hybrida Last, 1956a: 216 ( Zyras (Trachydonia), TL : Urundi, Rumonge, on the immediate banks of Lake

Tanganika, 800 m) Distribution: Burundi, Ghana, Nigeria, Tanzania, Zaire illotula Péringuey, 1904: 215 ( Myrmedonia, TL : Southern Rhodesia, Salisbury) Distribution: Rep. South Africa,

Zimbabwe, Uganda, Dem. Rep. Congo, Angola, new combination in Last, 1980b: 170 impudens Last, 1977c: 941 ( Zyras (Trachydonia), TL : Zaïre, Elisabethville) Distribution: Dem. Rep. Congo incontaminata Last, 1977c: 942 [= Last, 1967: 97, NOMEN NUDUM] ( Zyras (Trachydonia), TL : Katanga, reg.

Kolwezi) Distribution: Dem. Rep. Congo, Angola incorrupta Last, 1967: 111 ( Zyras (Trachydonia), TL : Angola, Marco de Canavezes, distr. Benguela) Angola,

Dem. Rep. Congo, Nigeria, Tanzania incredibilis Bernhauer, 1928a: 47 ( Zyras (Trachydonia), TL : Deutsch-Ostafrika, Kwakiyembe) Distribution:

Tanzania, Niger, Namibia, Kenya, Zaire, Rep. South Africa

= notabilis Pace, 1999b: 189 ( Diplopleurus, TL : Namibia, East Caprivi, Katima Mulilo) new synonymy incurva Last, 1956a: 217 ( Zyras (Trachydonia), TL : Ruanda, Mahembe, Nyanza territory, on the banks of River

Nyawarongo, 1400 m) Distribution: Rwanda kamerunensis Wendeler, 1930: 250 ( Zyras (Trachydonia), TL : Joko, Kamerun) Distribution: Cameroon klimai Bernhauer, 1929c: 190 ( Zyras (Trachydonia) ; TL: Kamerun, Joko) Distribution: Cameroon, Congo-

Brazzaville, Togo, Benin lathrimaeoides Bernhauer, 1908c: 116 ( Zyras, TL : Deutsh-Ostafrika, Dar el Salam) Distribution: Tanzania, Ghana,

Angola

= lathrimoeoides Bernhauer: Last, 1967: 96 (misspelling) levasseuri Last, 1966: 836 ( Zyras (Trachydonia), TL : Congo Republic, Odzala) Distribution: Congo-Brazzaville,

Dem. Rep. Congo, Ghana lucens Bernhauer, 1908c: 114 ( Zyras, TL : Deutsch-Ostafrika, Dar es Salam) Distribution: Congo-Brazzaville,

Tanzania, Uganda, Dem. Rep. Congo, Angola

= obscurior Last, 1977a: 192 ( Zyras (Trachydonia), TL : W. v. Albert-See, Awakubi a. Aruwimi) lusor Last, 1977c: 940 ( Zyras (Trachydonia), TL : Zaïre, Kivu, riv. Naturlonge, terr. Fizi) Distribution: Dem. Rep.

Congo, Tanzania majalis Bernhauer, 1928a: 51 ( Zyras (Trachydonia) lucens var. majalis, TL : Kwakiyeme) Distribution: Congo-

Brazzaville, Tanzania mandingorum Pace, 1988a: 27 ( Trachydonia, TL : Bissorã) Distribution: Guinea Bissau marginata Eppelsheim, 1885: 108 ( Myrmedonia, TL : Adda an der Goldküste [=Ada, Ghana]) Distribution: Ghana,

Cameroon, Angola methneri Bernhauer, 1908c: 116 ( Zyras, TL : Deutsch-Ostafrika, Usagara) Distribution: Tanzania, Ghana minarzi Bernhauer, 1929c: 194 ( Zyras (Trachydonia), TL : Deutsch-Ostafrika, Myombo) Distribution: Tanzania morogoroensis Last, 1981b: 116 ( Zyras (Trachydonia), TL : Tanzania, Morogoro) Distribution: Tanzania niziana Cameron, 1937b: 270 ( Zyras (Trachydonia), TL : Nizi, Blukwa) Distribution: Dem. Rep. Congo obnisa Last, 1967: 113 ( Zyras (Trachydonia), TL : Angola, Cameia) Distribution: Angola, Dem. Rep. Congo orientis Cameron, 1943b: 142 ( Zyras (Trachydonia) ; TL: W. Sarawak, Mt. Matang) Distribution: Malaysia

(Borneo: Sarawak) oxyteloides Bernhauer, 1908c: 115 ( Zyras, TL : Deutsch-Ostafrika, Dar es Salam) Distribution: Tanzania, Zambia,

Sudan, Dem. Rep. Congo pallescens Bernhauer, 1936f: 264 ( Zyras (Trachydonia) ; TL: Angola, Quirimbo) Distribution: Angola, Namibia,

Senegal panda Last, 1960d: 12 ( Zyras (Trachydonia), TL : Bogos) Distribution: Eritrea parcepunctata Bernhauer, 1929c: 195 ( Zyras (Trachydonia), TL : Deutsch-Ostafrika, Kilimandjaro-Gebiet)

Distribution: Tanzania, Ghana, Angola, Burundi, Rwanda, Dem. Rep. Congo

= sublucens Cameron, 1950b: 73 ( Zyras (Trachydonia), TL : Vitshumbi, S lac Édouard, 925 m) synonymy in

Last, 1967:96 patrizii Bernhauer, 1930a: 141 ( Zyras (Trachydonia), TL : Kongogebiet, Alto Uele, Fl. Duru) Distribution: Dem.

Rep. Congo peregrina Bernhauer, 1929c: 191 ( Zyras (Trachydonia), TL : Portugiesisch-Guinea, Bolama) Distribution: Guinea

Bissau, Congo-Brazzaville perinsignis Last, 1967: 116 ( Zyras (Trachydonia), TL : Angola, Réserve de Cameia) Distribution: Angola, Dem.

Rep. Congo persola Last, 1967: 114 ( Zyras (Trachydonia), TL : Angola, Réserve de Cameia) Distribution: Angola picea Bernhauer, 1928a: 44 ( Zyras (Trachydonia), TL : Deutsch-Ostafrika, Kilimandjaro-Gebiet) Distribution:

Tanzania piceola Cameron, 1950b: 73 ( Zyras (Trachydonia), TL : Kanyabayongo, Kabasha) Distribution: Dem. Rep. Congo privata Last, 1967: 110 ( Zyras (Trachydonia), TL : Angola, Réserve de Cameia, east of Vila Luso) Distribution:

Angola propinqua Bernhauer, 1928a: 43 ( Zyras (Trachydonia), TL : Deutsch-Ostafrika, Uha) Distribution: Tanzania,

Angola, Congo-Brazzaville, Dem. Rep. Congo, Rwanda puncticauda Bernhauer, 1930a: 142 ( Zyras (Trachydonia), TL : Belgisch-Kongo, Alto Uele, Kapili) Distribution:

Dem. Rep. Congo, Ghana, Congo-Brazzaville, Nigeria, Angola quadrimaculata Scheerpeltz, 1974a: 369 ( Zyras (Trachydonia), TL : Transvaal, Pretoria) Distribution: Rep. South

Africa rufoflava Cameron, 1939: 22 ( Zyras (Trachydonia), TL : Tjilatjap) Distribution: Indonesia (Java) sempiterna Last, 1967: 115 ( Zyras (Trachydonia), TL : Angola, Cameia) Distribution: Angola, Dem. Rep. Congo senator Last, 1967: 114 ( Zyras (Trachydonia), TL : Angola, Réserve de Cameia) Distribution: Angola signatipennis Last, 1981c: 3 [= Last, 1977a: 193, NOMEN NUDUM] ( Zyras (Trachydonia), TL : Okaunda, S. W.

Africa) Distribution: Namibia simulator Cameron, 1932: 39 ( Zyras (Trachydonia), TL : Haut Uele, Moto) Distribution: Dem. Rep. Congo sobria Last, 1967: 112 ( Zyras (Trachydonia), TL : Angola, Réserve de Cameia, east of Vila Luso) Distribution:

Angola sociabilis Last, 1967: 113 ( Zyras (Trachydonia), TL : Angola, Réserve de Cameia, east of Vila Luso) Distribution:

Angola solator Last, 1967: 115 ( Zyras (Trachydonia), TL : Angola, Cameia) Distribution: Angola speculator Last, 1967: 112 ( Zyras (Trachydonia), TL : Angola, Marco de Canavezes, distr. Benguela) Distribution:

Angola studiosa Last, 1967: 111 ( Zyras (Trachydonia), TL : Angola, Marco de Canavezes, distr. Bunguela) Distribution:

Angola, Ghana subasperata Bernhauer, 1929c: 192 ( Zyras (Trachydonia), TL : Portugiesisch Guinea, Bolama) Distribution:

Guinea Bissau, Congo-Brazzaville, Namibia subordinaria Last, 1967: 113 ( Zyras (Trachydonia), TL : Angola, Réserve de Cameia, east of Vila Luso)

Distribution: Angola taborae Bernhauer, 1928a: 49 ( Zyras (Trachydonia), TL : Deutsch-Ostafrika, Tabora) Distribution: Tanzania,

Angola thomasseti Bernhauer, 1928a: 45 ( Zyras (Trachydonia), TL : Natal, Weenen) Distribution: Rep. South Africa

(Natal), Guinea Bissau, Congo-Brazzaville, Dem. Rep. Congo, Tanzania, Uganda, Ghana trisulcus Bernhauer & Scheerpeltz, 1926: 706 ( Zyras , new name for Zyras trisulcatus Bernhauer, 1915 )

Distribution: Tanzania

= trisulcatus Bernhauer, 1915c: 171 ( Zyras, TL : Deutsch-Ostafrika, Daressalam, Westl.- Usambara;

preoccupied in Zyras , not Myrmedonia trisulcata Wasmann, 1896 ) tropica Kistner 2008b:505 (TL: Congo, Yangambi) Distribution: Angola, Cameroon, Gabon, Ghana, Uganda,

Dem. Rep. Congo, Zambia tuberculicollis Bernhauer, 1928a: 50 ( Zyras (Trachydonia), TL : Senegal) Distribution: Senegal ugandae Bernhauer, 1928a: 45 ( Zyras (Trachydonia), TL : Uganda, N. Chua) Distribution: Uganda uganoensis Bernhauer, 1947: 163 ( Zyras (Trachydonia), TL : Ugano) Distribution: Tanzania usagarae Bernhauer, 1928a: 42 ( Zyras (Trachydonia), TL : Deutsch-Ostafrika, S. O. Usagara) Distribution:

Tanzania, Angola vexans Cameron, 1948a: 243 ( Zyras (Trachydonia), TL : Moyen Chari, Fort Archambault) Distribution: Chad villiersi Cameron, 1952a: 834 ( Zyras (Trachydonia), TL : Togo, Klouto, Mt. Togo, 800 m) Distribution: Togo,

Angola vulnerata Bernhauer, 1928a: 43 ( Zyras (Trachydonia), TL : Deutsch-Ostafrika, Unteres Ruhembetal) Distribution:

Tanzania vulneriventris Bernhauer, 1928a: 44 ( Zyras (Trachydonia), TL : Sudan, Gell River Post, 70 m von Bahr-el-Gebel) Distribution: Sudan wichmanni Bernhauer, 1915c: 171 ( Zyras, TL : Abessinien, Harrar) Distribution: Ethiopia guara Last, 1980: 170 ( Zyras (Trachydonia), TL : not cited) Distribution: no locality cited, NOMEN NUDUM habitator Last, 1977b: 80 ( Trachydonia, TL : not cited) Distribution: Nigeria, NOMEN NUDUM titula Last, 1969: 145 ( Zyras (Trachydonia), TL : not cited) Distribution: Congo-Brazzaville, Tanzania, NOMEN NUDUM