Drusilla Leach, 1819

Hlaváč, Peter, Newton, Alfred F. & Maruyama, Munetoshi, 2011, World catalogue of the species of the tribe Lomechusini (Staphylinidae: Aleocharinae) 3075, Zootaxa 3075, pp. 1-151 : 32-34

Drusilla Leach

Drusilla Leach, 1819: 177 . Type species: Staphylinus canaliculatus Fabricius , by original designation

Drusilla Leach : Seevers, 1978: 152 (redescription); Maruyama, 2000 e: 351 (revision and key of Japanese species)

Drusilla Samouelle : Klimaszewski, 1982: 420 (redescription)

Astilbus Dillwyn, 1829:63 . New name for Drusilla Leach (unnecessary)

Astilbus Stephens : Cameron, 1939f: 460 (characters, key to species of India, Ceylon and Myanmar); Scheerpeltz, 1956 (key to palearctic species)

Agaricola Gistel, 1834: 10 . Type species: Staphylinus canaliculatus Fabricius , by monotypy

Myrmedonia Erichson, 1837: 286 . Type species: Staphylinus canaliculatus Fabricius , designated by Duponchel, 1841: 57

Camacopalpus Motschoulsky, 1858: 231 . Type species: Camacopalpus flavicornis Motschulsky , designated by Fenyes, 1918: 21; synonymy in Klimaszewski, 1982: 421.

Santhota Sharp, 1874: 3 . Type species: Santhota sparsa Sharp , by monotypy, synonymy in Maruyama, 2000 e: 351

Astilbus (Anaulax) Bernhauer, 1929d: 231 . Type species: Astilbus semicircularis Bernhauer , by monotypy; preoccupied, not de Roissy, 1805, not Murray, 1859; synonymy in Blackwelder, 1952: 137

Astilbus (Evansius) Bernhauer, 1933c: 297 . Type species: Astilbus denticollis Bernhauer , by monotypy

Thoracophagus Kistner, 2003d: 561 . Type species: Thoracophagus congoensis Kistner , by original designation; new synonymy subg. Apteranopsis Jeannel Apteranopsis Jeannel, 1960: 208. Type species: Apteranillus lethierryi Fauvel , by original designation Drusilla (Apteranopsis) Jeannel : Assing, 2002: 56 (subgenus of Drusilla ) Apteranillus Dohrn (misidentification): Fauvel, 1898 (key to species) convexifrons Fairmaire, 1873: 335 ( Apteranillus, TL : Dély-Ibrahim) Distribution: Algeria foreli demoflysi Normand, 1941: 174 ( Apteranillus, TL : Sidi-Tabet) Distribution: Tunisia foreli foreli Wasmann, 1890a: 318 [= Wasmann, 1890b: 319] ( Apteranillus, TL : Béja; HA: Aphaenogaster subterranea , A. testaceopilosa , A. crocea ) Distribution: Algeria, Tunisia hipponensis Fauvel, 1898: 108 ( Apteranillus, TL : Bône) Distribution: Algeria lethierryi Fauvel, 1898: 108 ( Apteranillus, TL : Province d’Alger, Hammam-Rhira, plateau entre Médéah et

Mouzaïa) Distribution: Algeria raffrayi Fairmaire, 1870: 371 ( Apteranillus, TL : Boghari) Distribution: Algeria theryi Fauvel, 1898: 109 ( Apteranillus, TL : Province de Constantine, col de Oliviers) Distribution: Algeria villosa Payerimhoff, 1909: 279 ( Apteranillus, TL : Fôret de Mizrana près Tigzirt, dép. d’Alger) Distribution:

Algeria subg. Drusilla Leach aino Nakane, 1963: 21 ( Drusilla, TL : Rausu, Hokkaido) Distribution: Japan (Hokkaidô, Honshû)

alutacea Reitter, 1901: 67 ( Drusilla (Astilbus), TL : Buchara: Karatak) Distribution: Iran, Turkmenistan, Uzbekistan, Tajikistan, Kazakhstan

= alutaceus Bernhauer, 1902a: 54 ( Astilbus, TL : Turkestan: Aulie Ata; preoccupied, not Drusilla (Astilbus) alutacea Reitter, 1901 ) synonymy in Bernhauer & Scheerpeltz 1926: 688

anceps Assing, 2005a: 130 ( Drusilla, TL : Turkey, Mersin, Sertavul Gecidi) Distribution: Turkey besucheti Assing, 2005a: 130 ( Drusilla, TL : Lebanon, Cèdres de Barouk) Distribution: Lebanon bulbata Assing, 2008c: 56 ( Drusilla, TL : Iran: Mazandaran, Elburz Mts., 20 km SSW Alamdeh, 36º22'N, 51º52'E, 1400m) Distribution: Iran canaliculata Fabricius, 1787: 221 ( Staphylinus, TL : Kiliae) Distribution: Europe, Russia (European, Siberia, Far East), Ukraine, Armenia, Georgia, Turkey, Azerbaijan, Israel, Iran, Kazakhstan; intro Canada (NB,ON,QC), USA (AK,KY,NY)

= aethiops Dalla Torre 1879: 52 ( Myrmedonia (Drusilla) canaliculata var. aethiops, TL : Oberösterreich)

= armeniacus Cameron, 1939b: 163 ( Astilbus canaliculatus armeniacus, TL : Armenia)

= caucasicus Bernhauer, 1903b: 186 ( Astilbus, TL : Caucasus) synonymy in Assing, 2005a: 113

= cavicollis Casey, 1906: 322 ( Drusilla, TL : Alaska)

= impressus Rossi, 1790: 254 ( Paederus, TL : provinciis Florentina et Pisana [ Italy])

= leonii Porta, 1907: 173 ( Astilbus, TL : Umbria, Abruzzo, Molise & Basilicata)

= mandli Bernhauer, 1899: 19 ( Astilbus, TL : Neusiedler See in Ungarn)

= puncticollis Motschulsky, 1845: 360 ( Drusilla, TL : Kamtschatka)

= sibiricus Scheerpeltz, 1956: 268 , 272 ( Astilbus, TL : Transbaikalien, Werchne-Udinsk) synonymy in Schilow, 1981b: 169

cernens Assing, 2005a: 125 ( Drusilla, TL : Syrie, Akbes) Distribution: Turkey, Israel, Syria cretica Assing, 2005a: 119 ( Drusilla, TL : Greece, Creta, Omalos) Distribution: Greece denigrata Assing, 2005a: 123 ( Drusilla, TL : Turkey, Antakya, 19 km south Antakya, south-west Şenköy)

Distribution: Turkey, Syria endorica Saulcy, 1864: 432 ( Myrmedonia, TL : Caifa, Syria [= Haifa, Israel]) Distribution: Israel, Syria;

synonymized by Gusarov, 1995: 85, revalidated by Assing, 2005a: 121, ganglbaueri Bernhauer, 1906: 127 ( Astilbus, TL : Turkestan, Mts. Karateghin, Sary-pul) Distribution: Iran,

Turkmenistan, Tajikistan, gracilis Hochhuth, 1849: 24 ( Myrmedonia, TL : Kaukasus) Distribution: Azerbaijan heydeni Eppelsheim, 1887: 313 ( Astilbus, TL : Kyndyr-Tau, Südwestl. Ausl. des Kara-Tau [ Uzbekistan])

Distribution: Iran, Uzbekistan, Turkmenistan, Kazakhstan, Kyrgyzstan, Tajikistan

= akinini Eppelsheim, 1888: 67 ( Astilbus, TL : Taschkent; HA: Crematogaster sp. , Tapinoma sp. , Lepisiota nigra ) synonymy in Assing, 2005b: 47. italica Bernhauer, 1903b: 186 ( Astilbus, TL : Vallo Lucania in Süditalien) Distribution: Italy kabyliana Fagel, 1958: 237 ( Astilbus, TL : Grand Kabylie, forêt d’Akfadou, fontaine des Houx) Distribution:

Algeria, Tunisia limata Assing, 2005a: 126 ( Drusilla, TL : Turkey, Antalya, Saklinet, 40 km west Antalya) Distribution: Turkey lydica Assing, 2007d: 36 ( Drusilla, TL : Aydin, 15 km NNE Aydin, Imambaba T., 1420 m) Distribution: Turkey memnonia Maerkel, 1844: 199 ( Astilbus, TL : Sicily; HA: Aphaenogaster subterranea ) Distribution: Italy (Sicily),

Algeria, Tunisia

= tristis Lucas, 1849: 99 ( Myrmedonia, TL : Moustapha-Supérieur & cimetière des Juifs, aux environs d'Alger)

synonymy in Assing, 2005a: 115 meridiana Fauvel, 1900: 244 ( Astilbus, TL : Morée: Cumani) Distribution: Greece (Pelopennese) nomurai Maruyama, 2000 e: 362 ( Drusilla, TL : Mt. Ôhoshi, Tsushima Is.) Distribution: Japan (Tsushima Is.) pallidicornis Assing, 2005a: 122 ( Drusilla, TL : Turkey, Antakya, Hatay, 2 km north Reyhauli, 50 km east Hatay)

Distribution: Turkey persica Assing, 2005a: 131 ( Drusilla, TL : Iran, Mazanderan, Tang-e-Râh) Distribution: Iran recta Assing, 2005a: 128 ( Drusilla, TL : Turkey, Mersin, road to Güzeloluk, S Aydinlar) Distribution: Turkey sinuosa Assing, 2005a: 127 ( Drusilla, TL : Turkey, Gaziantep, 33 km east Osmaniye, NE Nurdaği Geç)

Distribution: Turkey sparsa Sharp, 1874: 4 ( Santhota, TL : Nagasaki [LT; no locality cited by Sharp]) Distribution: Japan (Honshû,

Shikoku, Kyushu), South Korea taygetana Assing, 2005a: 118 ( Drusilla, TL : Greece, Taygetos, Morea) Distribution: Greece subg. Tropignorimus Bernhauer Astilbus (Tropignorimus) Bernhauer, 1915a: 154 . Type species: Astilbus carinithorax Bernhauer , by monotypy carinithorax Bernhauer, 1915a: 154 ( Astilbus (Tropignorimus) ; TL: Sarawak, Mt. Matang) Distribution: Malaysia

(Borneo: Sarawak) dusunorum Pace, 1993b: 162 ( Drusilla (Tropignorimus), TL : Sahah, Kinabalu N.P.) Distribution: Malaysia

(Borneo: Sabah) liangicola Pace, 1993b: 160 ( Drusilla (Tropignorimus), TL : S Celebes, Rante Pao) Distribution: Indonesia

(Sulawesi) manenekarum Pace, 1993b: 160 ( Drusilla (Tropignorimus), TL : S Celebes, Rante Pao) Distribution: Indonesia

(Sulawesi)