Lysilinga crassiseta Webb, 2006

Webb, Donald W. & Metz, Mark A., 2006, A Revision of the New World Genera Brachylinga Irwin and Lyneborg and Lysilinga Irwin and Lyneborg (Diptera: Therevidae: Therevinae) with the Description of a New Genus, Elcaribe Webb, Zootaxa 1288 (1288), pp. 1-241 : 218-220

Lysilinga crassiseta Webb spec. nov.

( Figs. 363, 373, 383, 393, 403, 413, 423, 433, 443, 449)

Etymology

Crassus (Latin) = thick, stout, seta (Latin) = bristle. This name is used as an adjective and refers to the large, stout, black macrosetae at the apex of the gonocoxite.

Diagnosis

Lysilinga crassiseta is taxonomically associated with L. pilifrons in possessing setae on the katepisternum and in possessing dorsocentral macrosetae. Lysilinga crassiseta differs from L. pilifrons in having the ventral setae on the forefemur white; anteroventral macrosetae present on the forefemur; the male tergite 8 lacking a basal sclerotized collar ( Fig. 363); the apex of the gonocoxite with two thick black macrosetae (Fig. 373); the subapical hood of the distiphallus large with the ventrolateral surface lacking a lobe (Fig. 433); and the female frons whitish brown dorsolateral to antennal base with a diffuse brown area dorsolaterally.

Description of holotype male (INBIO 165171)

Body length 6.7 mm.

Head. Head length 0.76 mm. Ommatidia smaller ventrally and laterally. Frons yellowish brown, surface raised; setae black, elongate. Antenna/head length 0.91; dark brown, pruinescence on scape and pedicel gray; scape length 0.24 mm, width 0.16 mm, length/width 1.5, scape/pedicel length 3.0, scape/flagellum width 1.0, setae black, elongate; pedicel length 0.08 mm, length/width 0.6; first flagellomere oval, attenuate apically, flagellum length 0.37 mm, 0.16 mm wide, length/width 2.3, flagellum/scape length 1.5. Parafacial pruinescence silver. Maxillary palpus orange brown, pruinescence gray; length 0.38 mm, length/width 5.4; setae white. Genal setae white. Occipital setae white becoming pale gold, lanceolate along margin of eye dorsally. Postocular macrosetae dark reddish brown.

Thorax. Macrosetae dark reddish brown, 3 np, 2 sa, 1 pa, 2 dc, 2 sc. Mesonotum brownish gray; setae black, elongate intermixed with gold, iridescent, lanceolate, appressed setae. Pleuron pruinescence gray; setae white on propleuron, anepisternum, katepisternum, laterotergite, metanepisternum, and scutellum. Wing. Membrane pale gray; veins brown; length 5.3 mm, length/width 3.1; pterostigma pale brown, narrow; cell m 3 closed at wing margin (closed, petiolate on right wing). Halter stalk dark brown, knob pale brown. Legs. Coxal macrosetae dark reddish brown. Femora dark reddish brown, pruinescence white; ventral setae white on fore­ and midfemur, dark reddish brown setae apicoventrally on forefemur; av 1:0:3, pv 0:0:2. Tibiae dark yellow, dark reddish brown apically particularly on hindtibia. Tarsomeres 1–2 dark yellow; 3–5 dark reddish brown.

Abdomen. Orange brown, pruinescence white, dense; dorsal setae white, appressed; lateral setae white, erect. Terminalia (INBIO 165174) dark yellowish brown. Tergite 8 and epandrium ( Fig. 363) quadrate, sides rounded; setae silver with scattered dark reddish brown setae. Hypoproct ( Fig. 363) large, posterior margin slightly emarginate; ending distal to apex of cercus. Gonocoxite ventral view (Fig. 373), basal half quadrate, median margin straight then curving apically to narrow apex, setae pale yellow basally becoming reddish brown apically with two thick, claw­like, apical macrosetae; dorsal view ( Fig. 383) gonocoxal bridge small; lateral view (Fig. 393), basal half broad, rounded, tapering apically to thick point; ventral lobe (Fig. 373) with ventral spines. Gonostylus ( Fig. 403), basodorsal lobe absent; ventral margin rounded, macrosetae dark yellow apically; fine, short spines apicoventrally. Aedeagus with dorsal apodeme (Fig. 413) broad, rectangular, posterolateral margin expanded; ventral apodeme ( Fig. 423) tapered anteriorly, extending to anterior margin of dorsal apodeme; distiphallus dorsal view (Fig. 413) tapered apically to point, lateral view (Fig. 433) broad basally, apical half directed ventrally then curving apically, subapical hood pointed, curved ventrally; ejaculatory apodeme (Fig. 413), anterior half greatly expanded laterally, extending anteriorly beyond anterior margin of dorsal apodeme, anterior margin truncate, in cross­section arched dorsoventrally with pair of dorsal carinas.

Variation (n=3). Body length 6.7–6.8, 6.7 mm. Thorax. Macrosetae 3–4 np, 2 sa, 1 pa, 2 dc, 2 sc. Legs. Anteroventral macrosetae 0–1:0:2–3, pv 0:0:1–2.

Female

Similar to male except for following.

Body length 8.8 mm.

Head. Frons surface raised, convex, brownish gray, whitish brown dorsolateral to antennal base with diffuse brown area dorsolaterally; setae dark brown over all of frons.

Thorax. Macrosetae 3 np, 2 sa, 1 pa, 2 dc, 2 sc. Legs. Anteroventral macrosetae 1:0:2, pv 0:0:1.

Abdomen. Dark reddish brown, glossy, posterolateral margin of tergites 2–6 with pruinescence whitish gray; dorsal setae dark reddish brown, short with pale gold, elongate, appressed setae along posterior margin of tergite 1; lateral setae dark reddish brown. Terminalia (INBIO 165173). Furca ( Fig. 443) oblong; length 0.48 mm, width 0.30 mm; posteromedial lobe short, conical; anterolateral projection short; anteromedial lobe absent. Common duct ( Fig. 443) longer than furca. Spermathecal duct ( Fig. 444) extended horizontally then directed anteriorly.

Distribution Lysilinga crassiseta is known from Costa Rica ( Fig. 449).

Habitats and phenology Adults of Lysilinga crassiseta have been collected in Malaise traps in February.

Specimens examined

Type specimen. The holotype male of Lysilinga crassiseta Webb (INBIO CR1002 165171) is labeled "San Luis, Monteverde, Prov. Punta. COSTA RICA, 1000–1350 m, Feb. 1995, Z. Fuentes, L_N_250850_449250, #4393" and is deposited in INBC. PARATYPES. COSTA RICA. GUANACASTE. Estación Cacao, 1100 m, 7–18.II.1995, R. H. Alvarado, 1♂ INB 196000 ( INBC), Malaise trap. PUNTARENAS. San Luis , Monteverde , 1000–1350 m, –. II.1995, Z. Fuentes, 1♂ INB 165174 ( INHS), 1♀ INB 165173 ( INBC), Malaise trap .