Titanophyllum spiliarum, Akkari, Nesrine, Stoev, Pavel & Enghoff, Henrik, 2011

Akkari, Nesrine, Stoev, Pavel & Enghoff, Henrik, 2011, Two new cave rnicolous genera of Julidae (Diplopoda, Julida), with notes on the tribe Brachyiulini and on julid subanal hooks and anchors, ZooKeys 114, pp. 1-14 : 3-4

Titanophyllum spiliarum sp. n. Figs 910−14

Material examined.

Holotype: adult ♂, Greece, Magnesia, Othris Mts., village of Kofi, Titanospilia (Cave of Titans), 13.VII.2003, P. Beron leg. (National Museum of Natural History Sofia − NMNHS); Paratypes: 4 ♂♂, 5 ♀♀, same locality, date and collector (ZMUC); 12 adult ♂♂, 11 adult ♀♀, 2 subadult ♂♂, 2 subadult ♀♀, same locality, date and collector (NMNHS).

Description of locality.

Titanospilia is an approximately 100 m long vertical cave composed of a single voluminous hall. All material was collected at the bottom of the shaft (P. Beron, pers. comm.).

Etymology.

The names means "of caves" in Greek and emphasizes the troglomorphic character of the species.

General description.

(all measurements in mm). Body uniformly pale to yellowish, legs brownish, metazonites with a slightly darker posterior band; length: 17.3−33.5 mm, vertical body diameter (height, H) 1-1.2 (♂) and 1-1.4 (♀); length/height ratio 16 (♂). Head: ocelli absent (Fig. 9); frons with 2 setae; gnathochilarium with 3 setae in apical part of each stipites, and with a long seta on each lamella lingualis; 4 supralabral setae and a row of ca 8 labral setae; mandibular stipites not expanded; male first leg-pair reduced and hook-shaped; antennal length about 1.6 ×H. Body rings with more or less dense striation and a whorl of moderately dense long setae; 50-61 (♂) and 47-54 (♀) podous rings, 1-2 apodous rings + telson. Defense glands visible as dark spots opening on the suture. Length of legs ca 0.83 ×H. Preanal ring dorsally only slightly protruding beyond anal margin; subanal scale with a small hook pointing anteriad (Fig. 14); anal valves pilose, with long setae. Male 7th body ring with well developed ventral lobes.

Gonopods.

Gonopods protruding from the 7th body ring. Anterior gonopod (promerite, p) much shorter than posterior gonopod (Figs 10, 11), uniformly broad along its length, slightly expanded at midlength; apex incised, with a lower angular process mesally and a higher triangular one laterally (Fig. 11). Posterior side of promerite at middle with a quite high, semitriangular ridge (r) pointing postero-laterad (Fig. 11). Flagellum (f) (Figs 10, 11) moderately long, falcate, pointing ventrad, emerging from the base of ridge.

Posterior gonopod (Figs 12, 13) unipartite, long, broad at base and midlength, gradually tapering distally; with a proximal lobe (lo) laterally (Fig. 12) and a subbasal fold (fo) mesally (Fig. 10), the latter giving rise to a mesal groove (g) running proximal and distal (Figs 10, 13) and ending in a subapical opening (op) (Fig. 12). Gonopodal apex (t) with a pointed tip (Figs 12, 13).