Nemastomatinae Simon, 1872a

SCHÖNHOFER, AXEL L., 2013, Axel L. Schönhofer, Zootaxa 3679 (1), pp. 1-68 : 26-29

Subfamily Nemastomatinae Simon, 1872a

Nemastomidae Simon, 1872a: 226 ; see Nemastomidae .

Nemastomatinae — Kratochvíl 1958: 530; type genus Nemastoma by subsequent designation (see Remarks); Gruber 1976: 800; Martens 1978: 93 (see Nemastomatidae ).

Giljaroviinae Kratochvíl, 1958: 530; type species Giljarovia rossica Kratochvíl, 1958 by original designation;—syn. Gruber 1976: 800.

Mitostominae Kratochvíl, 1958: 530; type species Phalangium chrysomelas Hermann, 1804 by original designation;—syn. Gruber 1976: 800.

Remarks. The genus Nemastoma was redefined to contain only a small set of mainly Central European species ( Gruber & Martens 1968), but up to now many other species of Nemastomatidae resided in their original placement as Nemastoma . Species are here pragmatically removed from Nemastoma sensu Gruber and Martens and assigned to other genera, while it is acknowledged that other authors were more careful in transferring species from Nemastoma to other genera (e.g. to Nemastomella ; Prieto 2004).

Kratochvíl (1958) investigated many of the species known in his days and established a widely accepted taxonomy. He unfortunately only provided species counts for his genera and rarely listed species names. Generic assignment is only attributed to Kratochvíl when clearly perceivable.

Genus Acromitostoma Roewer, 1951 (2 species)

Acromitostoma Roewer, 1951: 150 ; type species Acromitostoma rhinoceros Roewer, 1951 by monotypy and original designation;— Kratochvíl 1958: 530; Gruber 1976: 799; Rambla 1983: 22.

Carinostoma (Hispanostoma) Kratochvíl, 1958: 530 ; type species Carinostoma (Hispanostoma) hispanum Roewer, 1917 by monotypy;—syn. Gruber 1976: 799; Rambla 1983: 22.

Etymology. Gender neuter; combination of the Greek “akros” (pointed), referring to the long, spine-like tubercle on the eye mound of the type species, combined with the genus name „mitostoma“.

hispanum ( Roewer, 1917) ; loc. typ.: Spain, vicinity of Granada.

Nemastoma hispanum Roewer, 1917: 156 .

Carinostoma (Hispanostoma) hispanum — Kratochvíl 1958: 530.

Acromitostoma hispanum — Gruber 1976: 799; Rambla 1983: 22, 24 (redescr.).

Remarks. Gruber (1976: 799) synonymised Carinostoma (Hispanostoma) with Acromitostoma while Rambla (1983: 22) assumed he did not, yet she reconfirmed the synonymy.

rhinoceros ( Roewer, 1917) ; loc. typ.: Spain, near Malaga.

Nemastoma rhinoceros Roewer, 1917: 157 .

Acromitostoma rhinoceros — Roewer 1951: 150; Kratochvíl 1958: 530; Gruber 1976: 799.

Acromitostoma rhinocerus — Rambla 1983: 21 (misspelling, redescr.).

Genus Carinostoma Kratochvíl, 1958 (3 species)

Mitostoma (Carinostoma) Kratochvíl, 1958: 530 ; type species Nemastoma carinatum Roewer, 1914 by original designation. Carinostoma — Staręga 1976: 342; Martens 1978: 135.

Etymology. Gender neuter; combination of the Latin “carina” (keel), referring to the keels formed by anvil-shaped tubercles, and „stoma“, in reference to Nemastoma .

carinatum ( Roewer, 1914) ; loc. typ.: Herzegovina, Jablanica.

Nemastoma carinatum Roewer, 1914: 165 .

Mitostoma (Carinostoma) carinatum — Kratochvíl 1958: 530; Hadži 1973b: 14.

Carinostoma carinatum — Martens 1978: 135.

elegans (Sørensen, 1894) ; loc. typ.: Hungary (not further specified).

Nemastoma elegans Sørensen in Lendl, 1894: 29

Nemastoma elegans var. batorligetiense Szalay, 1952: 308 ;—syn. Martens 1978: 137.

Mitostoma (Carinostoma) elegans — Kratochvíl 1958: 572.

Carinostoma elegans — Martens 1978: 137.

Carinostoma elegans batorligetiense — Loksa 1991: 685.

Remarks. Loksa (1991: 685) re-established batorligetiense as a subspecies, which is not followed here as the outlined characters should be considered as variants.

ornatum ( Hadži, 1940) ; loc typ.: S Serbia (now Kosovo), Šar-Mountains, Kačanik.

Nemastoma ornatum Hadži, 1940: 8 .

Mitostoma (Mitostoma) ornatum — Hadži 1973b: 14.

Carinostoma ornatum — Staręga 1976: 342.

Remarks. When defining Carinostoma, Kratochvíl (1958) listed only carinatum and elegans . One can assume that he also included Nemastoma ornatum in Carinostoma as he mentioned three species in this well-defined group. Staręga (1976) formally placed ornatum in Carinostoma .

Genus Caucnemastoma Martens, 2006 (2 species)

Caucnemastoma Martens, 2006: 180 ; type species Caucnemastoma golovatchi Martens, 2006 by monotypy and original designation.

Etymology. Gender neuter; abbreviation of “Caucasus”, referring to the geographic origin, combined with “ Nemastoma ”.

golovatchi Martens, 2006: 180 ; loc. typ.: Russia, Krasnodar Prov., pasture Abago near Guzeripl. martensi Snegovaya, 2011 (in Snegovaya & Chumachenko 2011: 118); loc. typ.: Russia, Sochi, N Khosta,

Caucasian State Natural Biospheric Reserve, yew and box-tree grove.

Genus Centetostoma Kratochvíl, 1958 (4 species)

Centetostoma (Centetostoma) Kratochvíl, 1958: 530 ; type species Nemastoma centetes Simon, 1881b by monotypy and original designation;— Martens 1978: 139; 2011: 38.

Remarks. Without providing species names, Martens (1978: 140) included Nemastoma of the bacilliferum group discussed by Dresco (1967b) and Rambla (1968) within Centetostoma . Staręga (1986: 303) assigned the older available name Nemastomella to this assembly, gave a full list of species (1986: 304) but mentioned centetes as relatively isolated. Prieto (2004: 108) resolved this heterogeneity by restoring Centetostoma for centetes and Martens (2011: 38) finally redefined Centetostoma , including centetes and three species previously in Nemastoma .

Etymology. Gender neuter; combination of probably Greek “kentetes” (piercer), repeating and referencing the type species’ name (having rows of long, spine-like cones on the dorsal side) and „stoma“, in reference to Nemastoma .

centetes ( Simon, 1881b) ; loc. typ.: France, Dép. Alpes-Maritimes, St. Martin Lantosque.

Nemastoma centetes Simon, 1881b: 89 ;— Roewer 1923: 674.

Centetostoma centetes — Kratochvíl 1958: 530; Martens 1978: 140 (redescr.); 2011: 38; Prieto 2004: 108. Mitostoma centetes — Marcellino 1975: 124.

Nemastomella centetes — Staręga 1986: 304.

juberthiei Martens, 2011: 47; loc. typ.: France, Dép. Pyrénées-Orientales, SW Quillan, Lac des Bouillouses.

scabriculum ( Simon, 1879a) ; loc. typ.: France, Dép. Hautes-Pyrenées, Saint-Sauveur.

Nemastoma scabriculum Simon, 1879a: 284 ;— Rambla 1980: 198.

Centetostoma scabriculum — Martens 2011: 39 (redescr.).

ventalloi ( Mello-Leitao, 1936) ; loc. typ.: Spain, Lleida Prov., Val d’Aran.

Nemastoma ventalloi Mello-Leitao, 1936: 9 .

Centetostoma ventalloi — Martens 2011: 41 (redescr.).

Genus Giljarovia Kratochvíl, 1958 (10 species)

Giljarovia Kratochvíl, 1958: 530 ; type species Giljarovia rossica Kratochvíl, 1958 by monotypy;— Kratochvíl 1959: 1348; Gruber 1976: 797; Martens 2006: 151.

Malekia Kratochvíl, 1958: 530 ; type species Malekia stridula Kratochvíl, 1958 by monotypy;—syn. Gruber 1976: 797.

Remarks. Kratochvíl (1958) introduced and briefly defined Giljarovia and Malekia as new genera in 1958, but gave full descriptions in 1959, which was erroneously cited as the year of description by Martens (2006). Kratochvíl (1958) summarised both genera as Giljaroviinae on the basis of their unique stridulatory mechanism. Gruber (1976) rejected the subfamily and synonymised Malekia with Giljarovia , by claiming that stridulatory organs are not relevant at genus level.

Etymology. Gender feminine, as inferred from the feminine ending of species described together with the genus; in honour of the Russian arachnologist M.S. Giljarov.

kratochvili Snegovaya, 2011 (in Snegovaya & Chumachenko 2011: 119); loc. typ.: Russia, Sochi, N Khosta,

Caucasian State Natural Biospheric Reserve, yew and box-tree grove.

redikorzevi ( Charitonov, 1946) ; loc. typ.: Georgia, Sataplia Cave near Kutaissi.

Nemastoma redikorzevi Charitonov, 1946: 145 ;— Roewer 1951: 125.

Malekia redikorzevi — Staręga 1978: 201.

Giljarovia redikorzevi — Martens 2006: 165 (redescr.).

rossica Kratochvíl, 1958 ; loc. typ.: Russia, Krasnodar Prov., Kotch Mt. near Gorjacij Kljuc; Giljarovia rossica

Kratochvíl, 1958: 530;— Kratochvíl 1959: 1344; Martens 2006: 167 (redescr.).

stridula ( Kratochvíl, 1958) ; loc. typ.: Russia, Krasnodar Prov., Krasnaja Poljana.

Malekia stridula Kratochvíl, 1958: 530 ;— Kratochvíl 1959: 1348.

Giljarovia stridula — Gruber 1976: 798; Martens 2006: 151 (redescr.).

tenebricosa ( Redikortsev, 1936) ; loc. typ.: Russia, Suchumi Distr., upper Tsebelda Valley.

Nemastoma tenebricosum Redikortsev, 1936: 34 ;— Staręga 1978, 204.

Giljarovia tenebricosa — Martens 2006: 152 (redescr.). Remarks. Martens (2006: 155) proposed an isolated position within the genus as this species is missing a stridulatory apparatus.

thoracocornuta Martens, 2006: 167; loc. typ.: Russia, Daghestan, upper Gunib.

triangula Martens, 2006: 156; loc. typ.: Georgia, Caucasus Major, Kazbegi.

trianguloides Martens, 2006: 159; loc. typ.: Russia, Krasnodar Prov., Caucasian State Reserve, Pslukh, ca. 20 km E of Krasnaya Polyana.

turcica Gruber, 1976: 782; loc. typ.: Turkey, Vilayet Amasya , S Amasya.

vestita Martens, 2006: 161; loc. typ.: Russia, Krasnodar Prov., Caucasian State Reserve, Pslukh, ca. 20 km E of

Krasnaya Polyana, Mt. Kogot.