Ischyropsalidinae Simon, 1879a

SCHÖNHOFER, AXEL L., 2013, Axel L. Schönhofer, Zootaxa 3679 (1), pp. 1-68 : 12-16

publication ID

https://doi.org/ 10.11646/zootaxa.3679.1.1

publication LSID

lsid:zoobank.org:pub:770B1AA2-869F-4F1B-8010-0E3FAECB0592

persistent identifier

https://treatment.plazi.org/id/7E65412B-FE53-FFD5-FF2C-FC63E3F6FCE5

treatment provided by

Felipe

scientific name

Ischyropsalidinae Simon, 1879a
status

 

Subfamily Ischyropsalidinae Simon, 1879a View in CoL

Ischyropsalididae Simon, 1879a View in CoL (see there).

Ischyropsalidinae View in CoL ; new rank (compr. Ischyropsalis View in CoL ).

Diagnosis. Dyspnoi of the family Ischyropsalididae with a western Palearctic distribution. Defined by peculiarities of male genital morphology and enlarged chelicerae with glandular fields situated distally on the basal segment in males of most species. Palps long and thin, elongated to enable interaction with the chelicerae. Chelicerae massive and enlarged, longer than the body; in most species glandular fields of males associated with knob-like apophyses exposing the field to dorsal and/or distal side. Truncus penis as described for Ischyropsalididae ; glans penis usually simple and conical, but inflated and distinct from the truncus in many species; the sclerotised ventral plate mostly short, from roughly rhombic to deeply bi-lobed at the base; sclerotised area of glans with several defined regions exhibiting dense fields of mostly very thick, long and backward-pointing hairs; glans gradually tapering into a long stylus, the latter simple and pointed, bent at the base but without contortion or additional structures. Tuber oculorum without spine. Dorsal armature mostly lacking, dorsum smooth, often glossy, in few species opisthosomal areas raised to low, pointed bumps. Two or many spine-like metapeltidial structures, arranged in a transversal row.

Genus Ischyropsalis C.L. Koch, 1839b (22 species, 4 subspecies)

Ischyropsalis C.L. Koch, 1839b: 23 View in CoL ; type species Ischyropsalis kollari C.L. Koch, 1839b View in CoL by subsequent designation ( Thorell

1876: 467). Lhermia Lucas 1866: 44 View in CoL ; type species Lhermia spinipes Lucas 1866 View in CoL by monotypy;—syn. Simon 1972b: 484 (because found to be a nomen nudum); Martens 1969: 213. Ischiropsalis View in CoL — Simon 1872a: 227 (misspelling). Lermia View in CoL — Simon 1872b: 484 (misspelling). Ischyropsalis (Odontopalpa) Hadži, 1931: 113 ;—syn. Martens 1969: 189.

Remarks. Establishment of the present-day taxonomy and validating species-specific characters in Ischyropsalis have been mainly facilitated by the thorough revision of Martens (1969). Having been unable to borrow many types from the Iberian Peninsula, part of his work remained hypothetical for that local fauna, which was later corrected by Dresco (e.g. 1970, 1972), Luque (1991, 1992) and Prieto (1990a, 1990b). Of the 42 species described or emended by Roewer (1914, 1916, 1923, 1950, 1953a) only two remain valid. Species delineation and discovery of new species in northern Spain are still in progress (L. Labrada and C. Prieto, pers. comm.).

Etymology. Gender feminine; from Greek “ischyros” (robust, strong) and “psalis” (scissor), referring to the enlarged chelicerae.

adamii Canestrini, 1873 ; loc. typ.: Italian Peninsula, Golfo de Squillace.

Ischyropsalis adamii Canestrini, 1873: 50 View in CoL ;— Martens 1969: 205 (redescr.). Remarks. Martens (1969: 205) reported Roewer’s (1950: 33) lectotype of adamii View in CoL (SMF RI/976) as invalid and SMF data indicate that the lectotype locality could be “Como”, suggesting that the lectotype belongs to I. carli View in CoL . A lectoype designation is only necessary if the species definition is questionable (ICZN 75), which is currently not the case.

Ischyropsalis apuanus Caporiacco, 1930: 125 View in CoL ;—syn. Martens 1969: 205.

Ischyropsalis kratochvili Roewer, 1950: 32 View in CoL ;—syn. Martens 1969: 205.

Ischyropsalis amseli Roewer, 1950: 33 View in CoL ;—syn. Martens 1969: 205.

Ischyropsalis apuana — Roewer 1950: 36.

Ischyropsalis apuanus nanus Dresco, 1968b: 751 ; syn. n.: Dresco (1968b) described this subspecies, while Martens (1969) synonymised I. apuana likely without knowing about nanus.

cantabrica Luque & Labrada, 2012: 28; loc. typ.: Spain, Cantabria, Alfoz de Lloredo, Oreña Rogeria Cave.

carli Lessert, 1905 View in CoL ; loc. typ.: Switzerland, Graubünden. Remarks. Schuls-Fettan (today Scuol-Ftan), the first of three localities listed by Lessert (1905) and separated from the others by a distance of 40 km, is to be favoured as explicit type locality.

Ischyropsalis carli Lessert, 1905: 658 View in CoL .

Ischyropsalis pentelica Roewer, 1950: 30 View in CoL ;—syn. Martens 1969: 209.

dentipalpis Canestrini, 1872 View in CoL ; loc. typ.: Italy, Aosta Valley, Gressoney-St. Jean.

Ischyropsalis dentipalpis Canestrini, 1872: 9 View in CoL ;— Schönhofer & Martens 2010b: 4 (redescr.).

Ischyropsalis helvetica Roewer, 1916: 152 View in CoL ;—syn. Schönhofer & Martens 2010b: 4.

dispar Simon, 1872a View in CoL ; loc. typ.: Spain, Vizcaya, Orduño, Cueva de Albia.

Ischiropsalis dispar Simon, 1872a: 227 View in CoL .

Ischyropsalis goodnighti Roewer, 1950: 31 View in CoL ;—syn. Martens 1969: 256.

Ischyropsalis archeri Roewer, 1950: 48 View in CoL ;—syn. Martens 1969: 256.

Ischyropsalis noltei Dresco, 1972: 121 View in CoL ;—syn. Prieto 1990b: 286.

gigantea Dresco, 1968a: 308 View in CoL ; loc. typ.: Spain, Santander, Cueva la Cañuela.

hadzii Roewer, 1950 View in CoL ; loc. typ.: Slovenia, Mokritz Mountain, Ziavka Cave.

Ischyropsalis hadzii Roewer, 1950: 52 View in CoL ;— Martens 1969: 242 (redescr.). Remarks. Originally described as Phalangium cancroides Schmidt, 1851: 61 Roewer (1950) assumed the name preoccupied ( Müller 1776: 192) and changed it to hadzii View in CoL . Müller (1776) might have referred to Acarus cancroides Linnaeus, 1758: 616 (transferred to Phalangium View in CoL by Linnaeus 1767: 1028), now recognised as a pseudoscorpion. Meanwhile, the name hadzii View in CoL is in frequent use and established, and should not be altered. Roewer (1923: 683; also Hadži 1954: 141) questioned a specimen labelled “ Ischyropsalis cavernosum Schmidt ” (Krain, Zoological Institute University of Vienna) being the type of cancroides , which was not verified.

hellwigii View in CoL hellwigii ( Panzer, 1794) View in CoL ; loc. typ.: Germany, Elm Mountain (Elem) near Destedt, Braunschweig (reconstructed by Martens 1969). Remarks. Changes to the original spelling “ hellwigii View in CoL ” are prohibited (ICZN 33.4).

Phalangium hellwigii Panzer, 1794: 18 .

Ischyropsalis helwigii —C.L. Koch 1839b: 24.

Ischyropsalis herbstii C.L. Koch, 1847: 68 ;—syn. Martens 1969: 196.

Ischyropsalis hellwigii View in CoL —Sørensen in Lendl 1894: 20.

Ischyropsalis taunica Müller, 1924: 418 ;— Roewer 1923: 683 (as nomen nudum); syn. Martens 1965: 144; Martens 1969: 197. Remarks. Roewer pre-published a description provided by Müller and acknowledged Müller to be the author, who officially named it a year later. Hadži (1954) placed I. crassichelis and I. plicata in the synonymy of I. taunica , the latter synonymised by Martens (1965).

Ischyropsalis plicata Roewer, 1923: 692 ;— Hadži 1954: 159; syn. Martens 1969: 197.

Ischyropsalis pectinifera Hadži, 1928: 30 ;—syn. Hadži 1954: 159; Martens 1969: 197.

Ischyropsalis frankenbergeri Bartoš, 1938: 155 ;—syn. Martens 1969: 197.

Ischyropsalis franzi Roewer, 1950: 22 ;—syn. Martens 1969: 197.

Ischyropsalis crassichelis Roewer, 1950: 25 ;—syn. Hadži 1954: 159; Martens 1969: 197.

hellwigii View in CoL lucantei Simon, 1879a View in CoL ; loc. typ.: France, Dép. Hautes-Pyrénées, Grotte de Bétharram.

Ischyropsalis lucantei Simon, 1879a: 273 View in CoL ;— Dresco 1966: 814.

Ischyropsalis superbus Simon, 1881a: 129 View in CoL ;—syn. Martens 1969: 200.

Ischyropsalis superba — Simon 1881a: 130; Roewer 1914: 117.

Ischyropsalis hellwigii lucantei View in CoL — Martens 1969: 200. Remarks. Dresco (1966: 814) found lucantei View in CoL and superba conspecific but regarded the juvenile type of lucantei View in CoL as invalid, an error corrected by Martens (1969: 200).

hispanica Roewer, 1953a View in CoL ; loc. typ.: Spain, Sierra de Ancares, Cabana vella.

Ischyropsalis hispanica Roewer, 1953a: 208 View in CoL ;— Prieto 1990a: 319 (redescr.).

Ischyropsalis hispanum — Roewer 1953a: 208 (misspelling).

kollari C.L. Koch, 1839b View in CoL ; loc. typ.: Austria, Salzburg, Gastein.

Ischyropsalis kollari C.L. Koch, 1839b: 24 View in CoL —C.L. Koch 1840: 19 (description).

Ischyropsalis bosnica Roewer, 1914: 107 ;—syn. Martens 1969: 236.

Ischyropsalis (Odontopalpa) triglavensis Hadži, 1931: 113 ;—syn. Martens 1969: 236.

Ischyropsalis danubia Roewer, 1950: 21 ;— Hadži 1954: 152; syn. Martens 1969: 236.

Ischyropsalis strasseri Roewer, 1950: 22 ;—syn. Martens 1969: 236.

Ischyropsalis tirolensis Roewer, 1950: 23 ;—syn. Martens 1969: 236.

Ischyropsalis styriaca Roewer, 1950: 23 ;—syn. Martens 1969: 236.

Ischyropsalis reimoseri Roewer, 1950: 23 ;—syn. Martens 1969: 236.

Ischyropsalis spinichelis Roewer, 1950: 25 ;— Hadži 1954: 155; syn. Martens 1969: 236.

Ischyropsalis nivalis Roewer, 1950: 26 ;— Hadži 1954: 150; syn. Martens 1969: 236.

Ischyropsalis scutata Roewer, 1950: 26 ;— Hadži 1954: 156; syn. Martens 1969: 236.

Ischyropsalis segregata Roewer, 1950: 28 ;—syn. Martens 1969: 236.

Ischyropsalis troglodytes Roewer, 1950: 32 ;—syn. Martens 1969: 236.

lithoclasica Schönhofer & Martens, 2010b: 9 View in CoL ; loc. typ.: Italy, Bergamo, Stalle del Möschel.

luteipes Simon, 1872b View in CoL ; nomen protectum; loc. typ.: France, Ariège, Massat, grotte de Le Quère.

Lhermia spinipes Lucas 1866: 44 View in CoL ; nomen oblitum; Simon 1872b: 484. Remarks. Reviewing Lucas’ (1866) description, Lhermia spinipes View in CoL is presented in a sparse description, in a comparison with other harvestmen and with a distinct type locality. Simon’s (1872b) decision to treat the name as a nomen nudum and to provide the new name, Ischyropsalis luteipes View in CoL , is questionable, especially as he seems to have understood Lhermia spinipes View in CoL and I. luteipes View in CoL to be conspecific. However, it is proposed to maintain the use of luteipes View in CoL (ICZN 23.9.1; nomen protecum) because spinipes View in CoL (nomen oblitum) has not been used as a valid name after 1899 (ICZN 23.9.1.1) and luteipes View in CoL has been used in at least 60 works between 1962 and 2012 (ICZN 23.9.1.2).

Ischyropsalis luteipes Simon, 1872b: 484 View in CoL (replacement name).

Ischyropsalis pestae Roewer, 1950: 28 ;—syn. Martens 1969: 213.

Ischyropsalis caporiaccoi Roewer, 1950: 33 ;—syn. Martens 1969: 213.

Ischyropsalis caporioccoi — Martens 1969: 259 (misspelling).

Remarks. Dresco (1966: 598, 1970: 1205) defined three different forms or ecological races (intermedia, longipes , brevipes). Nothing is said about how to unambiguously recognise these forms, and they are of no nomenclatural relevance (ICZN 15.2).

magdalenae Simon, 1881a ; loc. typ.: Spain, Galmádes, Cueva de la Magdalena.

Ischyropsalis magdalenae Simon, 1881a: 130 View in CoL ;— Prieto 1990b: 289.

Ischyropsalis madalenae — Roewer 1914: 118 (misspelling).

manicata L. Koch, 1869 View in CoL ; loc. typ.: Romania (“Siebenbürgen”; not further specified).

Ischyropsalis manicata L. Koch, 1869: 164 View in CoL .

Ischyropsalis ianotai / ianotae Wajgel 1874: 36 (alternative spellings). Remarks. Wajgel (1874) mentioned the name ianotae in Ischyropsalis View in CoL , and Rafalski (1961: 15, then spelled ianotai ) recognised it as a nomen nudum.

Ischiropsalis manicatus — Lebert 1875 (incorrect ending).

Ischyropsalis dacica Roewer, 1916: 153 View in CoL ;—syn. Staręga 1966b: 148.

Ischyropsalis helvetica milleri Kratochvíl, 1934: 13 ;—syn. Martens 1969: 217.

Ischyropsalis kästneri Roewer, 1950: 19 View in CoL ;—syn. Martens 1969: 217.

Ischyropsalis austriaca Roewer, 1950: 22 View in CoL ;—syn. Martens 1969: 217.

Ischyropsalis balcanica Roewer, 1950: 30 View in CoL ;—syn. Martens 1969: 217.

Ischyropsalis milleri — Hadži 1954: 193.

Remarks. Martens (1969) states 1865 as the publication date which is related to a typo on his personal copy. Later authors have adopted this error.

muellneri Hamann, 1898 ; loc. typ.: Slovenia, Častitljiva Cave.

Ischyropsalis müllneri Hamann, 1898: 535 — Roewer 1950: 26.

Ischyropsalis mülleri — Hansen & Sørensen 1904: 5 (misspelling).

Ischyropsalis corcyraea Roewer, 1914: 108 ;—syn. Martens 1969: 245.

Ischyropsalis alfkeni Roewer, 1950: 19 ;—syn. Martens 1969: 245.

Ischyropsalis muellneri View in CoL — Martens 1965: 144; Martens 1969: 246 (redescr.).

navarrensis Roewer, 1950 View in CoL ; loc. typ.: Spain, Navarra. Remarks. Prieto (1990b) specified nicaea and turki as synonyms for having false type localities.

Ischyropsalis turki Roewer, 1950: 31 ;—syn. Prieto 1990b: 291.

Ischyropsalis nicaea Roewer, 1950: 36 ;—syn. Prieto 1990b: 291.

Ischyropsalis navarrensis Roewer, 1950: 44 View in CoL .

Ischyropsalis espagnoli Dresco, 1968c: 962 ;—syn. Prieto 1990b: 291.

nodifera Simon, 1879a View in CoL ; loc. typ.: France, Basses-Pyrénées, St. Jean de Luz.

Ischyropsalis nodifera Simon, 1879a: 270 View in CoL ;— Luque 1991: 28 (see Remarks on petiginosa View in CoL ).

Ischyropsalis sharpi Simon, 1879c: 129 View in CoL ;—syn. Simon 1881a: 128 (see Remarks on petiginosa View in CoL ).

Ischyropsalis janetscheki Roewer, 1950: 22 ;—syn. Martens 1969: 222; Prieto 1990a: 319.

Ischyropsalis moreana Roewer, 1950: 31 ;—syn. Martens 1969: 222; Prieto 1990a: 319.

Ischyropsalis corsica Roewer, 1950: 34 ;—syn. Martens 1969: 222; Prieto 1990a: 319.

Ischyropsalis asturica Roewer, 1950: 48 ;—syn. Martens 1969: 222; Prieto 1990a: 319.

petiginosa Simon, 1913 View in CoL ; loc. typ.: Spain, Oviedo, Pimiango, Première grotte de Mazaculos.

Ischyropsalis petiginosa Simon, 1913: 384 View in CoL ;— Luque 1991: 28.

Ischyropsalis pectiginosa — Mello-Leitao 1936: 12 (misspelling).

Remarks. Luque (1991) confirmed and redescribed nodifera and petiginosa , but did not unravel Martens’ synonymies (1969: 222; petiginosa and sharpi in syn. of nodifera ). A revision of these Ischyropsalis species needs to comment on the validity of petiginosa and sharpi .

pyrenaea pyrenaea Simon, 1872b ; loc. typ.: France, Ariège, Moulis, grotte de l’Estelas.

Ischiropsalis helwigi var. pyrenaea Simon, 1872b: 483 .

Ischyropsalis pyrenaea View in CoL — Bedel & Simon 1875: 128.

Ischyropsalis moreana Roewer, 1950: 31 ;—syn. Martens 1969: 228.

Ischyropsalis pyrenaea venasquensis Dresco, 1967a: 670 ;—syn. Martens 1969: 228.

Ischyropsalis pyrenaea pyrenaea View in CoL — Martens 1969: 227.

pyrenaea View in CoL alpinula Martens, 1978: 224; loc. typ.: Italy, Crissolo, Grotta di Rio Martino.

ravasinii Hadži, 1942: 11; loc. typ.: Italy, Cansiglio, Bus del Pal.

robusta Simon, 1872a View in CoL ; loc. typ.: Spain, Tras-os-Montes, Serra de Gerêz.

Ischiropsalis robusta Simon, 1872a: 230 View in CoL .

Ischyropsalis robusta View in CoL — Simon 1879a: 276.

Ischyropsalis lusitanica Roewer, 1923: 692 View in CoL ;—syn. Martens 1969: 233.

strandi Kratochvíl, 1936 View in CoL ; loc. typ.: Italy, Monte Baldo, Malgre Prato Cave.

Ischyropsalis strandi Kratochvíl, 1936: 248 View in CoL .

Ischyropsalis ruffoi Caporiacco, 1947: 139 View in CoL ;—syn. Juberthie 1963: 330.

Ischyropsalis knirschi Roewer, 1950: 27 ;—syn. Martens 1969: 249.

Kingdom

Animalia

Phylum

Arthropoda

Class

Arachnida

Order

Opiliones

Family

Ischyropsalididae

Loc

Ischyropsalidinae Simon, 1879a

SCHÖNHOFER, AXEL L. 2013
2013
Loc

lithoclasica Schönhofer & Martens, 2010b: 9

Schonhofer, A. L. & Martens, J. 2010: 9
2010
Loc

pyrenaea

Martens, J. 1978: 224
1978
Loc

Ischyropsalis noltei

Prieto, C. E. 1990: 286
Dresco, E. 1972: 121
1972
Loc

Ischyropsalis hellwigii lucantei

Martens, J. 1969: 200
Martens, J. 1969: 200
Dresco, E. 1966: 814
1969
Loc

Ischyropsalis caporioccoi

Martens, J. 1969: 259
1969
Loc

Ischyropsalis pyrenaea pyrenaea

Martens, J. 1969: 227
1969
Loc

Ischyropsalis apuanus nanus

Dresco, E. 1968: 751
1968
Loc

gigantea

Dresco, E. 1968: 308
1968
Loc

Ischyropsalis espagnoli

Prieto, C. E. 1990: 291
Dresco, E. 1968: 962
1968
Loc

Ischyropsalis pyrenaea venasquensis

Martens, J. 1969: 228
Dresco, E. 1967: 670
1967
Loc

Ischyropsalis muellneri

Martens, J. 1969: 246
Martens, J. 1965: 144
1965
Loc

Ischyropsalis milleri

Hadzi, J. 1954: 193
1954
Loc

Ischyropsalis hispanica

Prieto, C. E. 1990: 319
Roewer, C. F. 1953: 208
1953
Loc

Ischyropsalis hispanum

Roewer, C. F. 1953: 208
1953
Loc

Ischyropsalis kratochvili

Martens, J. 1969: 205
Roewer, C. F. 1950: 32
1950
Loc

Ischyropsalis amseli

Martens, J. 1969: 205
Roewer, C. F. 1950: 33
1950
Loc

Ischyropsalis apuana

Roewer, C. F. 1950: 36
1950
Loc

Ischyropsalis pentelica

Martens, J. 1969: 209
Roewer, C. F. 1950: 30
1950
Loc

Ischyropsalis goodnighti

Martens, J. 1969: 256
Roewer, C. F. 1950: 31
1950
Loc

Ischyropsalis archeri

Martens, J. 1969: 256
Roewer, C. F. 1950: 48
1950
Loc

Ischyropsalis hadzii

Martens, J. 1969: 242
Hadzi, J. 1954: 141
Roewer, C. F. 1950: 52
Roewer, C. F. 1950: 61
Roewer, C. F. 1923: 683
Muller, O. F. 1776: 192
Linnaeus, C. 1767: 1028
Linnaeus, C. 1758: 616
1950
Loc

Ischyropsalis franzi

Martens, J. 1969: 197
Roewer, C. F. 1950: 22
1950
Loc

Ischyropsalis crassichelis

Martens, J. 1969: 197
Hadzi, J. 1954: 159
Roewer, C. F. 1950: 25
1950
Loc

Ischyropsalis danubia

Martens, J. 1969: 236
Hadzi, J. 1954: 152
Roewer, C. F. 1950: 21
1950
Loc

Ischyropsalis strasseri

Martens, J. 1969: 236
Roewer, C. F. 1950: 22
1950
Loc

Ischyropsalis tirolensis

Martens, J. 1969: 236
Roewer, C. F. 1950: 23
1950
Loc

Ischyropsalis styriaca

Martens, J. 1969: 236
Roewer, C. F. 1950: 23
1950
Loc

Ischyropsalis reimoseri

Martens, J. 1969: 236
Roewer, C. F. 1950: 23
1950
Loc

Ischyropsalis spinichelis

Martens, J. 1969: 236
Hadzi, J. 1954: 155
Roewer, C. F. 1950: 25
1950
Loc

Ischyropsalis nivalis

Martens, J. 1969: 236
Hadzi, J. 1954: 150
Roewer, C. F. 1950: 26
1950
Loc

Ischyropsalis scutata

Martens, J. 1969: 236
Hadzi, J. 1954: 156
Roewer, C. F. 1950: 26
1950
Loc

Ischyropsalis segregata

Martens, J. 1969: 236
Roewer, C. F. 1950: 28
1950
Loc

Ischyropsalis troglodytes

Martens, J. 1969: 236
Roewer, C. F. 1950: 32
1950
Loc

Ischyropsalis pestae

Martens, J. 1969: 213
Roewer, C. F. 1950: 28
1950
Loc

Ischyropsalis caporiaccoi

Martens, J. 1969: 213
Roewer, C. F. 1950: 33
1950
Loc

Ischyropsalis kästneri

Martens, J. 1969: 217
Roewer, C. F. 1950: 19
1950
Loc

Ischyropsalis austriaca

Martens, J. 1969: 217
Roewer, C. F. 1950: 22
1950
Loc

Ischyropsalis balcanica

Martens, J. 1969: 217
Roewer, C. F. 1950: 30
1950
Loc

Ischyropsalis alfkeni

Martens, J. 1969: 245
Roewer, C. F. 1950: 19
1950
Loc

Ischyropsalis turki

Prieto, C. E. 1990: 291
Roewer, C. F. 1950: 31
1950
Loc

Ischyropsalis nicaea

Prieto, C. E. 1990: 291
Roewer, C. F. 1950: 36
1950
Loc

Ischyropsalis navarrensis

Roewer, C. F. 1950: 44
1950
Loc

Ischyropsalis janetscheki

Prieto, C. E. 1990: 319
Martens, J. 1969: 222
Roewer, C. F. 1950: 22
1950
Loc

Ischyropsalis moreana

Prieto, C. E. 1990: 319
Martens, J. 1969: 222
Roewer, C. F. 1950: 31
1950
Loc

Ischyropsalis corsica

Prieto, C. E. 1990: 319
Martens, J. 1969: 222
Roewer, C. F. 1950: 34
1950
Loc

Ischyropsalis asturica

Prieto, C. E. 1990: 319
Martens, J. 1969: 222
Roewer, C. F. 1950: 48
1950
Loc

Ischyropsalis moreana

Martens, J. 1969: 228
Roewer, C. F. 1950: 31
1950
Loc

Ischyropsalis knirschi

Martens, J. 1969: 249
Roewer, C. F. 1950: 27
1950
Loc

Ischyropsalis ruffoi

Juberthie, C. 1963: 330
Caporiacco, L. di 1947: 139
1947
Loc

Ischyropsalis frankenbergeri Bartoš, 1938: 155

Martens, J. 1969: 197
Bartos, E. 1938: 155
1938
Loc

Ischyropsalis pectiginosa

Mello-Leitao, C. F. de 1936: 12
1936
Loc

Ischyropsalis strandi Kratochvíl, 1936: 248

Kratochvil, J. 1936: 248
1936
Loc

Ischyropsalis helvetica milleri Kratochvíl, 1934: 13

Martens, J. 1969: 217
Kratochvil, J. 1934: 13
1934
Loc

Ischyropsalis (Odontopalpa) triglavensis Hadži, 1931: 113

Martens, J. 1969: 236
Hadzi, J. 1931: 113
1931
Loc

Ischyropsalis apuanus

Martens, J. 1969: 205
Caporiacco, L. di 1930: 125
1930
Loc

Ischyropsalis pectinifera Hadži, 1928: 30

Martens, J. 1969: 197
Hadzi, J. 1954: 159
Hadzi, J. 1928: 30
1928
Loc

Ischyropsalis taunica Müller, 1924: 418

Martens, J. 1969: 197
Martens, J. 1965: 144
Muller, A. 1924: 418
Roewer, C. F. 1923: 683
1924
Loc

Ischyropsalis plicata

Martens, J. 1969: 197
Hadzi, J. 1954: 159
Roewer, C. F. 1923: 692
1923
Loc

Ischyropsalis lusitanica

Martens, J. 1969: 233
Roewer, C. F. 1923: 692
1923
Loc

Ischyropsalis helvetica

Schonhofer, A. L. & Martens, J. 2010: 4
Roewer, C. F. 1916: 152
1916
Loc

Ischyropsalis dacica

Starega, W. 1966: 148
Roewer, C. F. 1916: 153
1916
Loc

Ischyropsalis bosnica

Martens, J. 1969: 236
Roewer, C. F. 1914: 107
1914
Loc

Ischyropsalis madalenae

Roewer, C. F. 1914: 118
1914
Loc

Ischyropsalis corcyraea

Martens, J. 1969: 245
Roewer, C. F. 1914: 108
1914
Loc

Ischyropsalis petiginosa

Luque, C. G. 1991: 28
Simon, E. 1913: 384
1913
Loc

Ischyropsalis carli

Lessert, R. de 1905: 658
1905
Loc

Ischyropsalis mülleri

Hansen, H. J. & Sorensen, W. 1904: 5
1904
Loc

Ischyropsalis müllneri

Roewer, C. F. 1950: 26
Hamann, O. 1898: 535
1898
Loc

Lendl, A. 1894: 20
1894
Loc

Ischyropsalis superbus

Martens, J. 1969: 200
Simon, E. 1881: 129
1881
Loc

Ischyropsalis superba

Roewer, C. F. 1914: 117
Simon, E. 1881: 130
1881
Loc

Ischyropsalis magdalenae

Prieto, C. E. 1990: 289
Simon, E. 1881: 130
1881
Loc

Ischyropsalis lucantei

Dresco, E. 1966: 814
Simon, E. 1879: 273
1879
Loc

Ischyropsalis nodifera

Luque, C. G. 1991: 28
Simon, E. 1879: 270
1879
Loc

Ischyropsalis sharpi

Simon, E. 1881: 128
Simon, E. 1879: 129
1879
Loc

Ischyropsalis robusta

Simon, E. 1879: 276
1879
Loc

Ischyropsalis pyrenaea

Bedel, L. & Simon, E. 1875: 128
1875
Loc

Ischyropsalis ianotai

Rafalski, J. 1961: 15
Wajgel, L. 1874: 36
1874
Loc

Ischyropsalis adamii

Martens, J. 1969: 205
Martens, J. 1969: 205
Roewer, C. F. 1950: 33
Canestrini, G. 1873: 50
1873
Loc

Ischyropsalis dentipalpis

Schonhofer, A. L. & Martens, J. 2010: 4
Canestrini, G. 1872: 9
1872
Loc

Ischiropsalis dispar

Simon, E. 1872: 227
1872
Loc

Ischyropsalis luteipes

Simon, E. 1872: 484
1872
Loc

Ischiropsalis helwigi var. pyrenaea

Simon, E. 1872: 483
1872
Loc

Ischiropsalis robusta

Simon, E. 1872: 230
1872
Loc

Ischyropsalis manicata L. Koch, 1869: 164

Koch, L. 1869: 164
1869
Loc

Lhermia spinipes

Simon, E. 1872: 484
Lucas, P. H. 1866: 44
1866
Loc

Ischyropsalis herbstii C.L. Koch, 1847: 68

Martens, J. 1969: 196
Koch, C. L. 1847: 68
1847
Loc

Ischyropsalis C.L. Koch, 1839b: 23

Koch, C. L. 1839: 23
1839
Loc

Ischyropsalis helwigii

Koch, C. L. 1839: 24
1839
Loc

Ischyropsalis kollari C.L. Koch, 1839b: 24

Koch, C. L. 1840: 19
Koch, C. L. 1839: 24
1839
Loc

Phalangium hellwigii

Panzer, G. W. F. 1794: 18
1794
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