Anaclastoctedon prionton, Plant, 2010

Plant, Adrian R., 2010, Anaclastoctedon (Diptera: Empididae: Hemerodromiinae), A New Genus From Asia And Australia, Raffles Bulletin of Zoology 58 (1), pp. 15-25 : 21-22

Anaclastoctedon prionton , new species

( Figs. 11, 12 )

Material examined. – Holotype. Male , AUSTRALIA: ACT, Canberra, Black Mountain, CSIRO, 35° 16'S, 149° 06'E, 20 Nov.–6 Dec.1998, coll. G. Gibson YPT ( CNC).

Paratypes. – Same data as holotype: 1 male; 1 male, 22–29 Nov. –1998, coll. G. Gibson MT ; 1 female 2–8 Nov.1998 (all in CNC) .

Diagnosis. – Black species, similar to A. ancistrodes new species, distinguished primarily by having wings yellowish, halter pale and distinctive male genitalia.

Description. – Male. Length 2.5mm. Head moderately dorsoventrally compressed, black, dusted greyish; face yellowish. Ocellar, vertical and upper postocular setae black, strong; Lower postocular setae yellowish, strong (particularly below), with weaker row in front close to eye margin. Proboscis yellowish brown, palpus pale with black seta apically. Antenna with scape black, pedicel yellowish brown or black, postpedicel black; stylus 5× long as postpedicel.

Thorax black, dusted greyish, setae blackish.

Legs rather pale yellow, distal tarsomeres darker, all setae black. F 1 very stout; about 3–5 av and 5–6 pv strong black setae between which are rows of about 14–15 av and 10–12

pv denticles. F 1 and F 3 with distinct dorsal fringe of setae. T 2 with strong apicoventral seta.

Abdomen brownish yellow, sternites 4–6 sometimes darker. Tergite 5 with fan of strong dark setae. Epandrium and hypandrium rather paler than rest of abdomen, bearing distinct setae; parameral sheath darker, black apically; cercus yellow. Epandrium subcircular, smaller than hypandrium ( Fig. 11 ). Cercus with major lobe gradually broadened distally, 6–7 stout black spines apically; basal lobe smaller, narrow. Parameral sheath ( Figs. 11, 12 ) long, conspicuously free for most of its length, subapically with curving flattened plate-like lateral process from which projects dorsal recurved sharply pointed process; apical process almost detached from basal part, minutely serrate dorsally about base, broadening at tip.

Wing membrane faintly yellowish, veins brownish-yellow; vein M 2 completely absent. Halter yellowish white.

Female. Similar to male but abdomen darker brown, postpedicel darker and apicoventral seta on T 2 shorter. F 1 with denticles rather more numerous, about 15 in pv row.

Etymology. – The specific epithet prionton (Greek) means serrate and refers to serrate apical process of the parameral sheath.

Remarks – This species is known only from Black Mountain in Australian Capital Territory in Eastern Australia during November and December. The habitat at the type locality is apparently dry sclerophyll forest (G. Gibson via B. Sinclair, pers. com.).

Figures

Figs. 9–12. Anaclastoctedon new genus: 9. A. ancistrodes new species, male terminalia; 10. A. sano new species, male terminalia; 11–12. A. prionton new species; 11. male terminalia; 12. apex of parameral sheath, dorsal aspect.