Ganesella hariola (Benson, 1856)

Sutcharit, Chirasak, Backeljau, Thierry & Panha, Somsak, 2019, Re-description of the type species of the genera Ganesella Blanford, 1863 and Globotrochus Haas, 1935; with description of a new Ganesella species from Thailand (Eupulmonata, Camaenidae), ZooKeys 870, pp. 51-76 : 51

Ganesella hariola (Benson, 1856) Figs 2 C–E , 8

Helix hariola Benson, 1856: 251. Pfeiffer 1860: 123, pl. 36, figs 21, 22. Hanley and Theobald 1870: 7, pl. 14, fig. 6.

Helix capitium var. hariola : Tryon 1887: 74, pl. 14, fig. 100.

Helix (Ganesella) hariola var. carinata Godwin-Austen, 1888: 242. Type locality: Khagan on Irrawaddy, and Hlindet, 1200 feet.

Ganesella capitium var. hariola : Pilsbry 1895: 170. Gude 1914: 197.

Type locality.

Thyet-Myo, prope ripas Irawadi fluvii [near the banks of the River Irrawaddy in Thayetmyo, Magway Region, Myanmar].

Material examined.

Type specimens. To stabilize the name, the syntype from Benson’s collection that most closely matched with the features and measurements of the original description is here designated as the lectotype UMZC I.104370/1 ( Fig. 2C , height 11.6 mm, width 14.4 mm) of Helix hariola Benson, 1856. The other shells from the same lot hence become the paralectotypes UMZC I.104370/2-4 (3 shells; Fig. 2D , height 13.8 mm, width 16.2 mm).

Additional material.

MYANMAR: Thungadan, North Ava, Burma: syntype of Helix hariola var. carinata NHMUK 1906.2.2.176 (4 shells). Burma: NHMUK 1906.2.2.276. Pegu: NHMUK Salisbury coll. ex. Beddome (1 shell). North Chin Hills, Upper Burma: NHMUK 1893.12.6.30-4 (5 shells). Thyet-myo, Pegu, Burma: NHMUK 1906.2.2.109 (4 shells). Pegu, Burma: ZMB Paetel coll. (2 shells), Dunker coll. (1 shell). Pyintha, Mandalay, Burma: ZMB Notling coll. (3 shells). Popa View Point Resort, about 50 km southeast of Bagan (20°55'19.1"N, 95°12'41.9"E), Kyaukpandaung Township, Nyaung-U District, Mandalay Region: CUMZ 5134 (1 shell; Fig. 2E ).

Description.

Shell small, dextral, thin and with a depressed trochoid. Apex acute; embryonic shell smooth with brownish colour. Whorls 5-6, increasing regularly, convex; suture wide and deep. Shell surface smooth or with fine growth lines. Last whorl large, very weekly angular to rounded; beneath convex; last whorl with brownish peripheral band. Shell brownish and translucent, with whitish-opaque, irregular streaks on upper and lower periphery. Aperture ovate; lip whitish and expanded; parietal callus transparent. Umbilicus rimate; columella whitish and expanded, overhanging umbilicus.

Remarks.

The shells of Ganesella hariola , G. carinella and G. capitium have a trochoid shape, but the shell of G. capitium is clearly more elevated than the shells of the two other species. The shell of G. hariola differs further from that of G. capitium and G. carinella in having a deep suture and a rounded last whorl with a brownish spiral band on the periphery. In contrast, the shells of G. capitium and G. carinella have a shallow suture and an angular to keeled last whorl without a brownish peripheral band.

Figures

Figure 2. A, B Ganesella capitium A lectotype UMZC I. 102385 / 1 and B paralectotype UMZC I. 102385 / 2 - 3 C-E Ganesella hariola C lectotype UMZC I. 104370 / 1 D paralectotype UMZC I. 104370 / 2 - 4 and E shell from Popa Mountain, Myanmar CUMZ 5134 F-H Ganesella carinella F lectotype SMF 27534 a G paralectotype SMF 27534 b and H shell from Takhli, Nakhonsawan CUMZ 5133.

Figure 8. Geographic distribution of Ganesella spp. and Globotrochus spp.