Stenus (Nestus) canaliculatus Gyllenhal, 1827

Ryvkin, Alexandr B., 2012, New species and records of Stenus (Nestus) of the canaliculatus group, with the erection of a new species group (Insecta: Coleoptera: Staphylinidae: Steninae), European Journal of Taxonomy 13, pp. 1-62 : 21-26

Stenus (Nestus) canaliculatus Gyllenhal, 1827

( Fig. 3D, E )

Stenus canaliculatus Gyllenhal, 1827: 501 .

Stenus canaliculatus – Fauvel 1869: 492. — J. Sahlberg 1871: 415. — Hochhuth 1872: 154. — Fauvel 1873: 255. —J. Sahlberg 1876: 55, 323. — Eppelsheim 1880: 94. — J. Sahlberg 1880: 77. — Heyden 1881: 78. — Casey 1884: 115. — Fauvel 1889: 119. — Hamilton 1889: 114. — Eppelsheim 1893: 56. — Hamilton 1894a: 20. — Hamilton 1894b: 371. — Poppius 1899: 38. — J. Sahlberg 1900: 28. — Fauvel 1902: 75. — Jakovlev 1902: 112. — L. Benick 1921a: 148. — L. Benick 1924: 254. — Lebedev 1925: 134. — Iljin 1926: 225. — Szujecki 1958: 118. — Puthz 1970b: 174, 178. — Korge 1971: 6. — Puthz 1971c: 139. — Puthz 1973a: 27. — Uhlig & Vogel 1981: 98. — Tichomirova 1982: 213. — Dubeshko 1984: 52. — Silfverberg 1988: 20 <‘holotype’ of congener Maeklin, 1853 >. — Pisanenko & Puthz 1991: 170, 174. — Puthz & Zanetti 1995: 17. — Shavrin 2007: 141. — Shavrin & Puthz 2007: 114 (pars).

Stenus (Nestus) canaliculatus – Rey 1884: 273. — Jakobson 1909: 481. — Poppius 1909a: 15. — Reitter 1909: 158. — Renkonen 1935: 29. — Palm 1961: 88. — Szujecki 1961: 34. — Puthz 1967b: 78. — Puthz 1971a: 82. — Puthz 1972a: 259. — Puthz 1972c: 12.— Tichomirova 1973: 173. — Shilov 1975: 58. — Ryvkin 1987: 159. — Ryvkin 1990: 168. — Kuznetsova 1991: 171. — Dauphin 1993: 188. — Ryabukhin 1999: 46. — Bordoni 2004: 120. — Semenov 2004: 11.

Stenus (s. str.) canaliculatus – Heyden 1898: 34. — Bernhauer & Schubert 1911: 156. — Scheerpeltz 1933: 1150. — Campbell & Davies 1991: 110. — Puthz 2001: 36.

Stenus affinis Stephens, 1833: 298 .

<?> Stenus congener Maeklin, 1853: 192 .

<?> Stenus congener – Casey 1884: 114. — Hamilton 1889: 114. — L. Benick 1921b: 114.

<?> Stenus (s. str.) congener – Scheerpeltz 1933: 1152.

Stenus cordaticollis Leinberg, 1900: 187 .

Stenus cordaticollis – J. Sahlberg 1900: 28.

Stenus (Nestus) cordaticollis – Jakobson 1909: 481.

Stenus (Nestus) foveifrons Rey, 1884: 272 .

Material examined

FRANCE: 1 ♀, ‘Caen (Ga) Raffray’ <white rectangle>, ‘Fvl[?Fauvel] Caen 67.’ <green circle>, ‘Fvl[?Fauvel] Stenus canaliculatus Gyll. ’ <white circle>, ‘c.[ollection] of A. Jakovlev’ <printed label> ( ZIN) .

POLAND: 1 ♂, ‘ Silesia. 4106[= Stenus canaliculatus Gyll. ] – 3’ ( ZIN) ; 1 ♀, ‘Danzig’, <underside> ‘Kraatz.’, ‘St. canaliculatus Gyll. ’, ‘c.[ollection] of A. Jakovlev’ <printed label> ( ZIN) .

HUNGARY: 3 ♂♂, 1 ♀, Nagycovaczy, 22.v.1982, V.G. Shilenkov leg. ( AR) ; 1 ♀, Budapest, 10 Apr. 1982, V.G. Shilenkov leg. ( ASh) .

UKRAINE: 1 ♀, Podol’sk Government, Kamenets-Podol’sk, 12 Apr. 1911, V. & I. Yakubovskiy leg. ( ZIN) ; 1 ♂, ‘ Odessa, Krivaya Balka . 4.iii.1920. D. Znojko leg.’, ‘ Stenus ( Nestus) morio Grav. ?’ ( ZIN) ; 1 ♀, Odessa, Fontanka , slope, lakeside, 4 Jul. 1970, S.Ya. Blinshtein leg. ( AR) ; 1 ♀, Nikolayev Area, Rybakovka , sandy seashore, 28 Jul. 1970, S.Ya. Blinshtein leg . ( AR) .

RUSSIA: 1 ♂, Kaliningrad Area, Rybachiy , 6 Jun. 1959, O.L. Kryzhanovskiy leg. ( ZIN) ; 1 ♀, same locality and collector, 28 Jun. 1959 ( ZIN) ; 1 ♂, Petropol .[is]. ( ZIN) ; 1 ♀, same locality, A. Moravitz Collection ( ZIN) ; 1 ♂, St. Petersburg, 30 Mar. 1897, Bianki leg., L. Benick det. ( ZIN) ; 1 ♂, 1 ♀, [St. Petersburg] Smolenskoye cemetery , 18 Aug. 1890 ( ZIN) ; 1 ♀, same locality, 5 Apr. 1891 ( ZIN) ; 1 ♀, same locality, 9 May 1896, G.G. Jakobson leg. ( ZIN) ; 2 ♂♂, 1 ♀, Petrograd [= St. Petersburg], Smolenskoye cemetery , 11 Apr. 1920, A.A. Stackelberg leg., L. Benick det. ( ZIN) ; 2 ♀♀, [St. Petersburg] Ekaterinhof , sifting, leaves, 2 Apr. 1921 ( ZIN) ; 1 ♀, ‘ Small Islands on Neva River , Shlisselburg Uyezd [=District], 1 Jun. 1906. G.G. Jakobson leg.’, ‘ canaliculatus ’ ( ZIN) ; 1 ♀, Pargolovo , Finlandskaya Railway , 8Aug. 1905, Somina leg., L. Benick det. ( ZIN) ; 1 ♂, St. Petersburg Government, Tsarskosel’skiy Uyezd [=District], Orlinskoye Lake , 26 Apr. 1905, D. Glazunov leg. ( ZIN) ; 1 ♀, [St. Petersburg Government], Luzhskiy District, Ploskoye , 7 Apr. 1906, Yu.I. Bekman leg. ( ZIN) ; 1 ♂, [? St. Petersburg Government], Lesnoy , 12 Oct. 1897 ( ZIN) ; 1 ♂, St. Petersburg Government, Luzhskiy Uyezd [=District], Preobrazhenskaya , 2 May 1905, D. Glazunov leg. ( ZIN) ; 1 ♀, ‘[St.-Petersburg Government,] Yamburg . [skiy] u.[yezd] [= Yamburgskiy District]. 12.1912.–01.1913. A.P. Rimskiy-Korsakov leg.’ ( ZIN) ; 1 ♀, Smolensk, bank of Dnepr River , on moist sand, 10 May 1993, M.Yu. Gildenkov leg. ( SSTC) ; 1 ♀, Moscow Government, Klinskiy Uyezd [=District], Boblovo , 16 Apr. 1906, D. Smirnov leg. ( ZIN) ; 1 ♀, Vladimir Area, Petushki , 1 May 1983, V.B. Semenov leg. ( AR) ; 1 ♀, same locality and collector, in litter, 21 Apr. 1984 ( AR) ; 1 ♀, Kostroma Area, Manturovskiy District, near Shilovo , Andromeda polifolia – Sphagnum magellanicum, Jun.1981 , A.L. Tichomirova leg. ( AR) ; 1 ♀, ‘Urzhum, drift of Urzhumka River, 21.iii.1901, [L. Krulikovskiy leg.]’, ‘ canaliculatus d. Bernh.[auer]’, ‘c.[ollection] of A. Jakovlev’ <printed label> ( ZIN) ; 1 ♀, ‘ Urzhum , drift of Urzhumka River . 23.iii.1901. [L. Krulikovskiy leg.]’, ‘ canaliculatus [?Bernhauer det.]’, ‘c.[ollection] of A. Jakovlev’ <printed label> ( ZIN) ; 1 ♀, ‘Urzhum , drift of Urzhumka River . 24–25.iii.1901. L. Krulikovskiy leg.’, ‘c.[ollection] of A. Jakovlev’ <printed label> ( ZIN) ; 1 ♂, 2 ♀♀, ‘[Urzhum,] drift of Urzhumka River . 09.iv.1906. L. Krulikovskiy leg.’, ‘ canaliculatus det. Shakhonin’, ‘c.[ollection] of A. Jakovlev’ <printed label> ( ZIN) ; 2 ♀♀, ‘ Malmyzh , drift of Shoshma River . 03–04.iv.1899. L. Krulikovskiy leg.’, ‘ canaliculatus ’, ‘c.[ollection] of A. Jakovlev’ <printed label> ( ZIN) ; 2 ♂♂, 3 ♀♀, ‘A. Jakowlew| Jaroslaw[l’ Government]’, ‘C.[ollection] of A. Jakovlev’, ‘ Stenus canaliculatus Gyll. d. Fauvel’, ‘c.[ollection] of A. Jakovlev’ <printed label> ( ZIN) ; 1 ♂, 1 ♀, ‘Yaroslav[l’]’, <underside> ‘Eppelsh[eim]’, ‘c.[ollection] of A. Jakovlev’ <printed label> ( ZIN) ; 1 ♂, ‘[Yaroslavl’ Government,] Manikha , sifting from floating litter. 15.iv.1898.’, ‘Yaroslavskiy U.[yezd] [=District].C.[ollection] of A.Jakowlew’, ‘ Stenus carbonarius Gyll. [det.? A.Jakowlew]’ ( ZIN) ; 1 ♀, Yaroslavl’ Area, Nekouzskiy District , Borok , shore of reservoir , 6 Jun. 1979, E.M. Veselova leg. ( AR) ; 1 ♀, ‘Ryaz[an]’<white rectangle with thin black frame and species name> ‘ Stenus aterrimus Er. ’, ‘C.[ollection] of A. Jakowlew’ ( ZIN) ; 1 ♂, 1 ♀, ‘Riasan.’ <printed label on red paper>, ‘ Stenus canaliculatus Gyll. ’ <black Indian ink in dubble box> ( ZIN: ex coll. Semenov-Tian-Shanskiy) ; 1 ♀, Gremyachka , Dankovskiy Uyezd [=District], Ryazan’ Government, 13 Aug. 1899, A. Semenov leg. ( ZIN: ex coll. Semenov-Tian-Shanskiy) ; 1 ♀, same locality and collector, 19 Sep. 1901 ( ZIN) ; 1 ♀, Tataria , Stolbishchevskiy District , collective farm ‘13 Let Oktyabrya’, perennial grass, clover: in trap buckets, 1 Jun. 1956, Utrobina leg. ( AR) ; 1 ♀, Voronezh Area, Khopyorskiy Nature Reserve , near water , 5 Jun.? 1976. ( AR) ; 1 ♂, Samara Area, Kinel’skiy District , Krasnosamarskoye Forestry , flood-plain of Samara River , 2 Jul. 1995, I.N.Goreslavets leg., ‘ Stenus sp-6 det. Goreslavetz I.N. ’ ( IG) ; 1 ♀, Saratov Area, Khvalynsk , Sterkh River , 30 Jul. 1974, V.I. Lomakin leg. ( AR) ; 1 ♀, [Rostov Area] Env. Novocherkassk , Razliv , 17 Apr. 1913, V. Kiseritzky leg. ( ZIN) ; 1 ♀, Rostov Area, Nizshne-Kundryuchenskoye hunting farm , 2 Jun. 1990, V. Grebennikov leg. ( ZMRU) ; 1 ♀, ‘Sarepta. [18]95. Becker.’, ‘ canaliculatus d. Bernh.[auer]’, ‘c.[ollection] of A. Jakovlev’ <printed label> ( ZIN) ; 3 ♂♂, Astrakhan’ Area, near Dosang , 16 May 1997, K.A. Grebennikov leg., ‘ Stenus (Nestus) ? canaliculatus Gyll. K.A. Grebennikov det.1997’ ( KG) ; 1 ♀, Kalmyk ASSR , Lysyi Liman , 26 Jun. 1977, Arzanov leg. ( ZMRU) ; 1 ♂, Kalmykiya, Arshanya Zelmen’ , 3 Jun. 1980, N.S. Kalyuzhnaya leg. ( KG) ; 1 ♀, NW Caucasus [Krasnodar Territory], Malyi Tkhach Mt. , 2000 m a.s.l., subalpine belt , clayey bank of puddle, 4 Jun. 1994, A.Yu. Solodovnikov leg. ( AYuS) ; 1 ♀, Murmansk Area, Poyakonda , Biological Field Research Station of Moscow University , ejected Fucus , 14 Jun. 1979, E.M. Veselova leg. ( AR) ; 1 ♀, same locality and collector, Alopecurus ventricosus , 13–17 Jul. 1979 ( AR) ; 1 ♀, near Solovetskiy Monastery , Bol’shoy Solovetskiy Island , 1893, G.G. Jakobson leg. ( ZIN) ; 1 ♂, Komi ASSR , Gibyu River , near Ukhta , fluvial terrace above flood-plain , in sod, 26 Jun. 1974, V.F. Shilov leg. ( ZIN) ; 1 ♂, Komi ASSR , near Ukhta , in drift after flood , 23 Jun. 1974, V.F. Shilov leg. ( ZIN) ; 1 ♂, same locality and collector, 5 May 1974 ( ZIN) ; 1 ♂, Sob’ River basin, Bol’shoy Ural, Obdorsk [=Salekhard], 8 Jul. 1925, Fridolin leg. ( ZIN) ; 1 ♀, Tyumen Area, Uvatskiy District , 10–11 km S of Gornoslinkino, steep right bank of Irtysh River up-stream of Tobolsk Field Research Station of Severtsov Institute, small mosses, plant debris, denuded loam among Tussilago farfara , Carex sp., Poaceae gen. sp., Fabaceae etc. near small rill draining by slope , 11 Jun. 2004, A.B. Ryvkin leg. ( AR) ; 1 ♀, Tyumen Area, Uvatskiy District , 11 km S of Gornoslinkino, clayish right bank of Irtysh River (narrow fold at rill mouth) upstream of Tobolsk Field Research Station of Severtsov Institute , in leaf litter and sweeping under Salix sp. with small true mosses, Marchantia sp., Equisetum sp., Carex sp., Calamagrostis sp. etc., 11 Jun. 2004, A.B. Ryvkin leg. ( AR) ; 1 ♂, 2 ♀♀, Tomsk Government , Ob’ River between Barnaul and Chesnokovka (postal station), 13 Jun. 1897, Silant’yev leg. ( ZIN) ; 1 ♂, 1 ♀, Tuva, East shore of Chagytay Lake , river bank , 18 Jul. 1980, B.A. Korotyayev leg. ( AR) ; 1 ♂, Krasnoyarsk Territory, Beryozovskiy District, ‘Stolby’ Nature Reserve , Mana River near Berly cordon , shingles at river bank, 22 Jun. 1990, A.B. Ryvkin leg. ( AR) ; 1 ♀, Evenkia , Baykitskiy District, Central Siberian Biosphere Reserve , Stolbovaya River basin : Dulkuma River (IV), 12 km below Topkaya River mouth, 165 m a.s.l., river banks : leaf litter among Poaceae gen. sp. and Salix sp., shingles, sand, silt, drift, 1 Jul. 1990, A.B.Ryvkin leg. ( AR) ; 1 ♂, Evenkia , Baykitskiy District , Central Siberian State Reserve , Stolbovaya River basin : lower reaches of Birapchana River near mouth of Kruten’kiy Stream , 110 m a.s.l., river bank : shingles, silt, sand, clay, Poaceae gen. sp., mosses, Carex spp., etc., 21 Jul. 1991, A.B. Ryvkin leg. ( AR) ; 2 ♂♂, Krasnoyarsk Territory, Turukhanskiy District , Nizhnyaya Sarchikha River near Kamenka River mouth , 150 m a.s.l., under mosses on river bank, 6–7 Jul. 1992, V.B. Semenov leg. ( AR) ; 1 ♀, same locality and collector, river bank : sand, 10 Jul. 1992 ( AR) ; 1 ♂, Krasnoyarsk Territory, Turukhanskiy District , Yeloguy River , 7–10 km below Tyna River mouth, 70 m a.s.l., sandy river bank with small shingles, 19 Jul. 1989, A.B. Ryvkin leg. ( AR) ; 1 ♂, 1 ♀, same locality and collector, river bank : Carex spp., Poaceae gen. sp., mosses, shingles, 24 Jul. 1989 ( AR) ; 1 ♂, same locality, Sphagnum bog with Eriophorum sp., Ledum palustre , Oxycoccus sp., Rubus chamaemorus, Vaccinium uliginosum and sparse Pinus sylvestris , 1 Jul. 1989, R.A. Rakitov leg. ( AR) ; 1 ♂, 3 ♀♀, Krasnoyarsk Territory, Turukhanskiy District , Yeloguy Refuge , Yeloguy River 6 km below Tyna River mouth, river bank : under drift, 22–23 Jul. 1992, V.B. Semenov leg. ( AR) ; 1 ♂, Krasnoyarsk Territory, Turukhanskiy District , Yeloguy Refuge , Tyna River 12 km up-stream of river mouth, under mosses on stony bank of river , 12 Aug. 1992, V.B. Semenov leg. ( AR) ; 1 ♂, Putorana Highland , Ayan Lake near Ayan River source , in stony stream channel , 9 Jul. 1983, K.Yu. Eskov leg. ( AR) ; 1 ♂, Taymyr, Maymecha River mouth , 5 Jul. 1971, A.P. Rasnitsyn, A.G. Ponomarenko, I.D. Sukatcheva, V.V. Zherikhin leg. ( AR) ; 1 ♂, Taymyr, Khatangskiy District , Kotuy River , 6 km up-stream of Kresty, Salix sp., 13 Jul. 1976, A.P. Rasnitsyn & I.D. Sukatcheva leg. ( AR) ; 1 ♂, 5 ♀♀, ‘ Irkutsk. V.Ye. Yakovlev leg.’, ‘ canaliculatus Gyll. L. Benick det.’ ( ZIN) ; 1 ♀, ‘ Irkutsk. V.Ye. Yakovlev leg.’, ‘ canal. v. cordaticollis Leinbg. L. Benick det.’ <immature specimen> ( ZIN) ; 1 ♂, ‘ Irkutsk. V.Ye. Yakovlev leg.’, ‘Coll. Semenov-Tian-Shansky’ ( ZIN) ; 1 ♂, Buryatia, Selenga River , sedge community, 3 Jul. 1982, Dubeshko leg. ( AR) ; 1 ♂, Buryatia, middle reaches of Vitim River , Baysa , 3 Aug. 1969, V.V. Zherikhin, Ye. Shnitnikova leg. ( AR) ; 1 ♂, [ Yakutia ], ‘5 verst off Yakutsk, suburban settlement. 18.vi.1925. Bianki leg.’, ‘Yakutian research expedition of Acad. of Sci.’ ( ZIN) ; 1 ♀, ‘ Yakutsk. 21.ix.1927. Moskvin leg.’, ‘168’, ‘ canaliculatus Gyll. L. Benick det.’ ( ZIN) ; 1 ♂, ‘2nd Tyhylyminskiy Nasleg, Yakutsk Region. 02.vii.1925. Bianki leg.’, ‘Yakutian research expedition of Acad. of Sci.’, ‘ canaliculatus Gyll. L. Benick det.’ ( ZIN) ; 1 ♀, Central Yakutia, Nizhniy Bestyakh ( ASh); 1 ♂, W Yakutia , 11 km SW of Toybokhoy, Uchugey Syhylakh, sedge-gramineous hygrophytous meadow, 25 Aug. 1997, Stepanov leg., ‘ Stenus carbonarius Gyll. ’ ( ASh) ; 1 ♀, Khabarovsk Territory, Okhotsk , Airport , wall of timber house, 4 Jul. 1985, V. Zherikhin, A. Rasnitsyn, D. Shcherbakov leg. ( AR) ; 1 ♀, Maritime Prov. , Khasan , under boards on littoral, 3 Jun. 1967, A.L. Tichomirova leg. <?> ( AR) .

MONGOLIA: 1 ♂, Seleng.[a] Aimak, Shamar, 18 Jul. 1984, S.L. Kuz’min leg. ( AR) .

USA: 1 ♀, ‘ Wisconsin: Walworth Co.; Troy Center. ( SW) Girl Scout Camp Area. W. Suter leg. 27: VI:1976.’, ‘FM( HD)#76-92. WS#76-55. mosses ex boggy field.’ ( FMNH) ; 1 ♂, ‘ USA: Ill., Kendall Co., Silver Springs St. Pk. 26-IV-1981.’, ‘ FMHD #81-164, ex rotting wood chips, L. Watrous leg.’ ( FMNH) ; 1 ♀, ‘ USA: Mich. Berrien Co., 2 mi. N. New Buffalo, Gallien R. W bank 2-xi-80.’, ‘litter in marsh. L.E. Watrous leg.’ ( FMNH) ; 1 ♀, ‘19’ <indian ink on white rectangle>, ‘Kenai’ <indian ink on green stripe>, ‘ Stenus | congener | Mäkl .| Am. b. occ.’ <indian ink on green rectangle> ( ZMMU: Motschulsky collection) .

Remarks

The species is variable in both external characters and, to a lesser extent, in the shape of the aedeagus. Terra typica: ‘Suecia’ [= Sweden]. Widespread in the Holarctic. In an earlier article devoted to the Caucasian stenine fauna, I summarized the range as follows: ‘Palearctic and Alaska <…>, Michigan <…>, but missing evidently in subtropics and in the southernmost districts of the Pontic biogeographical province (Turkestanian and Kurdian). Unknown to me also from the Crimean, Derbentian, Caucasian, Lenkoranian districts. The records nearest to the Caucasus are in the Rostov Area (Nedvigovka, Azhinov, Manychskaya, Yerokhin, Podtyolkovo, Liventsovka). One cannot deny a possibility of finding it in Ciscaucasia’ ( Ryvkin 1990; the biogeographical units follow those of Razumovskiy 1980). In this excerpt I missed the important record from NE Turkey: ‘Benliahmed südwestl. Kars’ ( Korge 1971); though the locality does not belong to the Caucasus in a strict sense, it lies right at the border between the Caucasian and Kurdian biogeographical districts. This species is missing in the lists for the NW Caucasus ( Solodovnikov 1998) and the Samara Area ( Goreslavets et al. 2002), but my restudy of the collections of the named authors makes it possible to report S. canaliculatus for both the regions (see above: the Material section).

Both records of S. canaliculatus in Shavrin (1998) (for “Alar', Irkutsk Area”, and “Bol'shoye Alginskoye Lake, Buryatia”) are to be referred to S. incrassatus Erichson, 1839 (I have seen the material in Shavrin’s collection). Among the material listed under the name ‘ canaliculatus ’ by Shavrin & Puthz (2007), at least the specimen from ‘Barun-Torej, Myrgen’, Chita Area should be referred to the new species S. canalis sp. nov. described below; some other Transbaikalian specimens unknown to me may also prove to belong to this new species. In that article, all the records marked as relating to the material deposited in ‘coll. A.Shavrin’ and ‘Irkutsk State University’, except ‘Sarma’, ‘Shelekhov’, and ‘Selenga delta’ (the latter locality, based on my identifications, has been published by Dubeshko (1984)), should be verified because the respective specimens were not present in the complete collection that Dr. Shavrin has sent me for restudy.

Ryabukhin (1999), based on my identifications in the IBPM collection, listed three localities from Kamchatka.

Campbell & Davies (1991) reported this species for Alaska and all the provinces of Canada, excluding Saskatchewan, Labrador, and Prince Edward Island, without citing material. The American specimens I know from the northernmost localities are very similar to those from NE Asia; but the material from the USA main territory as well as from S Canada demonstrates definite differences both in body proportions (e.g. the head somewhat more wide) and in the shape of the male genitalia; I believe the variability mentioned above requires further analysis based on more representative sampling.

S. congener Maeklin, 1853 , according to the original description, was described for a single specimen captured on Kadjak Island (‘In insula Kadjak a D.Holmberg semel captus’). No specimens from the Kenai Peninsula were mentioned in Maeklin’s text; these two localities were distinguished well by both Mäklin and Mannerheim (a good map was provided in Mannerheim 1853, which incorporated Maeklin’s descriptions); therefore, assumptions by Silfverberg (1988) and Puthz (2001) concerning the type status of Kenai specimens in different museums seem to be as premature as unfounded (‘semel captus’<!> means that the type series cannot be distributed widely over all the main European museums). Casey (1884), L. Benick (1921b), and Scheerpeltz (1933) treated S. congener as a species propria; nevertheless, Fauvel (1869, 1873, 1889), Rey (1884), Bernhauer & Shubert (1911), and others regarded this name as a synonym of S. canaliculatus . I think it important that Fauvel (1889), when citing the synonymy, mentioned Maeklin’s type as known to him and cited the terra typica as ‘Kadjak’. Consequently, the synonymy of congener and canaliculatus should be regarded as hypothetical before a reliable type specimen can be found.

Figures

Fig. 3. – A–C: Stenus (Nestus) alopex sp. nov.: ♂. A. Aedeagus, ventral view (holotype). B. Apical part of aedeagus, ventral view (holotype). C. Abdominal sternite 9 (paratype). – D–E: S. (N.) canaliculatus Gyllenhal, 1827: ♂. D. Aedeagus, ventral view (Russia: Rostov Area: Nedvigovka). E. Abdominal sternite 9 (Russia: Taimyr: Maymecha River). – Scales = 0.1 mm.